D. Sarris, D. Christodoulakis, and C. Körner, Recent decline in precipitation and tree growth in the eastern Mediterranean, Global Change Biology, vol.14, issue.6, pp.1187-1200, 2007.
DOI : 10.1111/j.1365-2486.1996.tb00054.x

L. Misson, D. Degueldre, C. C. Rodriguez, R. Rocheteau, A. Ourcival et al., Phenological responses to extreme droughts in a Mediterranean forest, Global Change Biology, vol.23, issue.100, pp.1036-1048, 2011.
DOI : 10.1111/j.1365-2486.2010.02348.x

W. Thuiller, S. Lavorel, M. Aráujo, M. Sykes, and I. Prentice, Climate change threats to plant diversity in Europe, Proceedings of the National Academy of Sciences, vol.102, issue.23, pp.8245-8250, 2005.
DOI : 10.1073/pnas.0409902102

URL : https://hal.archives-ouvertes.fr/halsde-00274241

J. Lenoir, J. Gégout, P. Marquet, P. De-ruffray, and H. Brisse, A Significant Upward Shift in Plant Species Optimum Elevation During the 20th Century, Science, vol.320, issue.5884, pp.1768-1771, 2008.
DOI : 10.1126/science.1156831

URL : https://hal.archives-ouvertes.fr/hal-01195113

L. Hughes, Biological consequences of global warming: is the signal already apparent?, Trends in Ecology & Evolution, vol.15, issue.2, pp.56-61, 2000.
DOI : 10.1016/S0169-5347(99)01764-4

Q. Thomas, R. Canham, C. Weathers, K. Goodale, and C. , Increased tree carbon storage in response to nitrogen deposition in the US, Nature Geoscience, vol.99, issue.1, pp.13-17, 2009.
DOI : 10.1038/ngeo721

N. Bréda, A. Granier, and G. Aussenac, Effects of thinning on soil and tree water relations, transpiration and growth in an oak forest (Quercus petraea (Matt.) Liebl.), Tree Physiology, vol.15, issue.5, pp.295-306, 1995.
DOI : 10.1093/treephys/15.5.295

N. Mcdowell, J. Brooks, S. Fitzgerald, and B. Bond, Carbon isotope discrimination and growth response of old Pinus ponderosa trees to stand density reductions, Plant, Cell and Environment, vol.282, issue.4, pp.631-644, 2003.
DOI : 10.1007/s004420000609

D. Martín-benito, D. Río, M. Heinrich, I. Helle, G. Cañellas et al., Response of climate-growth relationships and water use efficiency to thinning in a Pinus nigra afforestation, Forest Ecology and Management, vol.259, issue.5, pp.967-975, 2010.
DOI : 10.1016/j.foreco.2009.12.001

G. Gea-izquierdo, D. Martín-benito, P. Cherubini, and I. Cañellas, Variabilit?? des relations climatcroissance chez Quercus ilex L. dans des peuplements forestiers ouverts de diff??rentes densit??s dans l???ouest de la p??ninsule Ib??rique, Annals of Forest Science, vol.48, issue.17, pp.802-802, 2009.
DOI : 10.1051/forest/2009080

E. Piutti and A. Cescatti, A quantitative analysis of the interactions between climatic response and intraspecific competition in European beech, Canadian Journal of Forest Research, vol.27, issue.3, pp.277-284, 1997.
DOI : 10.1139/x96-176

D. Martín-benito, V. Kint, M. Del-río, B. Muys, and I. Cañellas, Growth responses of West-Mediterranean Pinus nigra to climate change are modulated by competition and productivity: Past trends and future perspectives, Forest Ecology and Management, vol.262, issue.6, pp.1030-1040, 2011.
DOI : 10.1016/j.foreco.2011.05.038

J. Martínez-vilalta, B. López, L. Loepfe, and F. Lloret, Stand- and tree-level determinants of the drought response of Scots pine radial growth, Oecologia, vol.32, issue.3, pp.877-888, 2012.
DOI : 10.1007/s00442-011-2132-8

E. Vaganov, M. Hughes, and A. Shashkin, Growth dynamics of conifer tree rings. Images of past and future environments, 2006.

G. Gea-izquierdo, L. Fernández-de-uña, and I. Cañellas, Growth projections reveal local vulnerability of Mediterranean oaks with rising temperatures, Forest Ecology and Management, vol.305, pp.282-293, 2013.
DOI : 10.1016/j.foreco.2013.05.058

C. Canham, M. Papaik, M. Uriarte, W. Mcwilliams, J. Jenkins et al., Neighborhood Analyses Of Canopy Tree Competition Along Environmental Gradients In New England Forests, Ecological Applications, vol.16, issue.2, pp.540-554, 2006.
DOI : 10.1139/cjfr-30-10-1571

A. Fichtner, K. Sturm, C. Rickert, W. Härdtle, and J. Schrautzer, L. varies with tree size and abiotic stress: minimizing anthropogenic disturbances in forests, Journal of Applied Ecology, vol.54, issue.6, pp.1306-1315, 2012.
DOI : 10.1111/j.1365-2664.2012.02196.x

URL : https://hal.archives-ouvertes.fr/halshs-00914552

J. Landsberg and P. Sands, Physiological ecology of forest production: principles, processes and models, 2011.

F. Ni, T. Cavazos, M. Hughes, A. Comrie, and G. Funkhouser, Cool-season precipitation in the southwestern USA sinceAD 1000: comparison of linear and nonlinear techniques for reconstruction, International Journal of Climatology, vol.10, issue.13, pp.1645-1662, 2002.
DOI : 10.1002/joc.804

S. Tolwinski-ward, M. Evans, M. Hughes, and K. Anchukaitis, An efficient forward model of the climate controls on interannual variation in tree-ring width, Climate Dynamics, vol.5, issue.1, pp.2419-2439, 2010.
DOI : 10.1007/s00382-010-0945-5

L. Andreu, E. Gutiérrez, M. Macias, M. Ribas, O. Bosch et al., Climate increases regional treegrowth variability in Iberian pine forests, Glob Chang Biol, vol.13, pp.804-815, 2007.

I. Climate-change-solomon, S. Qin, D. Manning, M. Chen, Z. Marquis et al., The Physical Science Basis Contribution of Working Group I to the Fourth Assessment Report of the Intergovernmental Panel on Climate Change, 2007.

D. Ruiz-labourdette, D. Nogués-bravo, H. Ollero, M. Schmitz, and F. Pineda, Forest composition in Mediterranean mountains is projected to shift along the entire elevational gradient under climate change, Journal of Biogeography, vol.66, issue.1, pp.162-176, 2012.
DOI : 10.1111/j.1365-2699.2011.02592.x

F. Múgica, F. García-antón, M. , S. Ollero, and H. , Vegetation dynamics and human impact in the Sierra de Guadarrama, Central System, Spain. The Holocene, pp.69-82, 1998.

R. Sánchez-de-dios, M. Benito-garzón, and H. Sainz-ollero, Present and future extension of the Iberian submediterranean territories as determined from the distribution of marcescent oaks, Plant Ecology, vol.41, issue.1, pp.189-205, 2009.
DOI : 10.1007/s11258-009-9584-5

A. Rigling, C. Bigler, B. Eilmann, E. Feldmeyer-christe, U. Gimmi et al., Driving factors of a vegetation shift from Scots pine to pubescent oak in dry Alpine forests, Global Change Biology, vol.63, issue.1, pp.229-240, 2013.
DOI : 10.1111/gcb.12038

M. Valbuena-carabaña, S. González-martínez, and L. Gil, Coppice forests and genetic diversity: A case study in Quercus pyrenaica Willd. from Central Spain, Forest Ecology and Management, vol.254, issue.2, pp.225-232, 2008.
DOI : 10.1016/j.foreco.2007.08.001

I. Cañellas, G. Montero, and Á. Bachiller, Transformation of quejigo oak (Quercus faginea Lam.) coppice forest into high forest by thinning, Ann Ist Sper Selvic, vol.27, pp.143-147, 1996.

I. Cañellas, D. Río, M. Roig, S. Montero, and G. , Willd. coppice stands in Spanish central mountain, Annals of Forest Science, vol.61, issue.3, pp.243-250, 2004.
DOI : 10.1051/forest:2004017

L. Corcuera, J. Camarero, and E. Gil-pelegrín, Functional groups in Quercus species derived from the analysis of pressure???volume curves, Trees, vol.16, issue.7, pp.465-472, 2002.
DOI : 10.1007/s00468-002-0187-1

B. Eilmann, R. Zweifel, N. Buchmann, P. Fonti, and A. Rigling, Drought-induced adaptation of the xylem in Scots pine and pubescent oak, Tree Physiology, vol.29, issue.8, pp.1011-1020, 2009.
DOI : 10.1093/treephys/tpp035

G. Montserrat-martí, J. Camarero, S. Palacio, C. Pérez-rontomé, R. Milla et al., Summer-drought constrains the phenology and growth of two coexisting Mediterranean oaks with contrasting leaf habit: implications for their persistence and reproduction, Trees, vol.5, issue.100, pp.787-799, 2009.
DOI : 10.1007/s00468-009-0320-5

R. Duursma, P. Kolari, M. Perämäki, E. Nikinmaa, P. Hari et al., Predicting the decline in daily maximum transpiration rate of two pine stands during drought based on constant minimum leaf water potential and plant hydraulic conductance, Tree Physiology, vol.28, issue.2, pp.265-276, 2008.
DOI : 10.1093/treephys/28.2.265

. Rinntech and . Tsap-win, Time series analysis and presentation for dendrochronology and related applications . Version 0.53, 2003.

R. Holmes, Computer assisted quality control in tree-ring dating and measurement. Tree-Ring Bull, pp.69-78, 1983.

F. Biondi and F. Qeadan, A Theory-Driven Approach to Tree-Ring Standardization: Defining the Biological Trend from Expected Basal Area Increment, Tree-Ring Research, vol.64, issue.2, pp.81-96, 2008.
DOI : 10.3959/2008-6.1

G. Gea-izquierdo and I. Cañellas, Analysis of holm oak intraspecific competition using Gamma regression, For Sci, vol.55, pp.310-322, 2009.

S. Herrera, J. Gutierrez, R. Ancell, M. Pons, M. Frias et al., Development and analysis of a 50-year high-resolution daily gridded precipitation dataset over Spain (Spain02), International Journal of Climatology, vol.5, issue.2, pp.74-85, 2012.
DOI : 10.1002/joc.2256

T. Mitchell and P. Jones, An improved method of constructing a database of monthly climate observations and associated high-resolution grids, International Journal of Climatology, vol.78, issue.6, pp.693-712, 2005.
DOI : 10.1002/joc.1181

M. Kirchner, T. Faus-kessler, G. Jakobi, M. Leuchner, L. Ries et al., Altitudinal temperature lapse rates in an Alpine valley: trends and the influence of season and weather patterns, International Journal of Climatology, vol.49, issue.2, pp.539-555, 2013.
DOI : 10.1002/joc.3444

W. Goffe, G. Ferrier, and R. J. , Global optimization of statistical functions with simulated annealing, Journal of Econometrics, vol.60, issue.1-2, pp.65-99, 1994.
DOI : 10.1016/0304-4076(94)90038-8

J. Johnson and K. Omland, Model selection in ecology and evolution, Trends in Ecology & Evolution, vol.19, issue.2, pp.101-108, 2004.
DOI : 10.1016/j.tree.2003.10.013

K. Burnham and D. Anderson, Model selection and multimodel inference: a practical information-theoretic approach, 2002.
DOI : 10.1007/b97636

H. Pretzsch and P. Biber, A re-evaluation of Reineke's rule and stand density index, For Sci, vol.51, pp.304-320, 2005.

R. Development and C. Team, R: A language and environment for statistical computing. R Foundation for Statistical Computing Available: http://www.r-project.org, 2011.

A. Bunn, A dendrochronology program library in R (dplR), Dendrochronologia, vol.26, issue.2, pp.115-124, 2008.
DOI : 10.1016/j.dendro.2008.01.002

L. Murphy and . Likelihood, Methods for maximum likelihood estimation. R package version 1.5 Available: http://cran.r-project.org/package = likelihood, 2012.

L. Gómez-aparicio, R. García-valdés, P. Ruíz-benito, and M. Zavala, Disentangling the relative importance of climate, size and competition on tree growth in Iberian forests: implications for forest management under global change, Global Change Biology, vol.196, issue.7, pp.2400-2414, 2011.
DOI : 10.1111/j.1365-2486.2011.02421.x

C. Moreno-gutiérrez, G. Battipaglia, P. Cherubini, M. Saurer, E. Nicolás et al., Stand structure modulates the long-term vulnerability of Pinus halepensis to climatic drought in a semiarid Mediterranean ecosystem, Plant, Cell & Environment, vol.23, issue.6, pp.1026-1039, 2012.
DOI : 10.1111/j.1365-3040.2011.02469.x

V. Rozas and I. García-gonzález, Too wet for oaks? Inter-tree competition and recent persistent wetness predispose oaks to rainfall-induced dieback in Atlantic rainy forest, Global and Planetary Change, vol.94, issue.95, pp.94-95, 2012.
DOI : 10.1016/j.gloplacha.2012.07.004

W. Webb, M. Newton, and D. Starr, Carbon dioxide exchange of Alnus rubra, Oecologia, vol.10, issue.4, pp.281-291, 1974.
DOI : 10.1007/BF00345747

G. Farquhar and T. Sharkey, Stomatal Conductance and Photosynthesis, Annual Review of Plant Physiology, vol.33, issue.1, pp.317-345, 1982.
DOI : 10.1146/annurev.pp.33.060182.001533

K. Hikosaka, K. Ishikawa, A. Borjigidai, O. Muller, and Y. Onoda, Temperature acclimation of photosynthesis: mechanisms involved in the changes in temperature dependence of photosynthetic rate, Journal of Experimental Botany, vol.57, issue.2, pp.291-302, 2006.
DOI : 10.1093/jxb/erj049

D. Arrigo, R. Wilson, R. Liepert, B. Cherubini, and P. , On the ???Divergence Problem??? in Northern Forests: A review of the tree-ring evidence and possible causes, Global and Planetary Change, vol.60, issue.3-4, pp.289-305, 2008.
DOI : 10.1016/j.gloplacha.2007.03.004

C. Loehle, A mathematical analysis of the divergence problem in dendroclimatology, Climatic Change, vol.112, issue.D11, pp.233-245, 2008.
DOI : 10.1007/s10584-008-9488-8

J. Vayreda, J. Martinez-vilalta, M. Gracia, and J. Retana, Recent climate changes interact with stand structure and management to determine changes in tree carbon stocks in Spanish forests, Global Change Biology, vol.329, issue.3, pp.1028-1041, 2012.
DOI : 10.1111/j.1365-2486.2011.02606.x

L. Galiano, J. Martínez-vilalta, and F. Lloret, Drought-Induced Multifactor Decline of Scots Pine in the Pyrenees and Potential Vegetation Change by the Expansion of Co-occurring Oak Species, Ecosystems, vol.72, issue.9, pp.978-991, 2010.
DOI : 10.1007/s10021-010-9368-8

C. Bigler, J. Gri?ar, H. Bugmann, and K. ?ufar, Growth patterns as indicators of impending tree death in silver fir, Forest Ecology and Management, vol.199, issue.2-3, pp.183-190, 2004.
DOI : 10.1016/j.foreco.2004.04.019

P. Chen, C. Welsh, and A. Hamann, Geographic variation in growth response of Douglas-fir to interannual climate variability and projected climate change, Global Change Biology, vol.55, issue.12, pp.3374-3385, 2010.
DOI : 10.1111/j.1365-2486.2010.02166.x

J. Huang, Y. Bergeron, F. Berninger, L. Zhai, J. Tardif et al., Impact of Future Climate on Radial Growth of Four Major Boreal Tree Species in the Eastern Canadian Boreal Forest, PLoS ONE, vol.35, issue.5, p.23468879, 2013.
DOI : 10.1371/journal.pone.0056758.s007

N. Bréda, R. Huc, A. Granier, and E. Dreyer, Temperate forest trees and stands under severe drought: a review of ecophysiological responses, adaptation processes and long-term consequences, Annals of Forest Science, vol.63, issue.6, pp.625-644, 2006.
DOI : 10.1051/forest:2006042

N. Mcdowell, W. Pockman, C. Allen, D. Breshears, N. Cobb et al., Mechanisms of plant survival and mortality during drought: why do some plants survive while others succumb to drought?, New Phytologist, vol.40, issue.4, pp.719-739, 2008.
DOI : 10.1007/BF00328910

G. Gea-izquierdo, B. Viguera, M. Cabrera, and I. Cañellas, Drought induced decline could portend widespread pine mortality at the xeric ecotone in managed mediterranean pine-oak woodlands, Forest Ecology and Management, vol.320, pp.70-82, 2014.
DOI : 10.1016/j.foreco.2014.02.025

J. Martínez-vilalta and J. Piñol, Drought-induced mortality and hydraulic architecture in pine populations of the NE Iberian Peninsula, Forest Ecology and Management, vol.161, issue.1-3, pp.247-256, 2002.
DOI : 10.1016/S0378-1127(01)00495-9

P. Ruiz-benito, E. Lines, L. Gómez-aparicio, M. Zavala, and D. Coomes, Patterns and Drivers of Tree Mortality in Iberian Forests: Climatic Effects Are Modified by Competition, PLoS ONE, vol.11, issue.2, p.23451096, 2013.
DOI : 10.1371/journal.pone.0056843.s008

R. Benavides, S. Rabasa, E. Granda, A. Escudero, J. Hódar et al., Direct and Indirect Effects of Climate on Demography and Early Growth of Pinus sylvestris at the Rear Edge: Changing Roles of Biotic and Abiotic Factors, PLoS ONE, vol.17, issue.8, p.23555794, 2013.
DOI : 10.1371/journal.pone.0059824.s002

Y. Kouba, J. Camarero, and C. Alados, Roles of land-use and climate change on the establishment and regeneration dynamics of Mediterranean semi-deciduous oak forests, Forest Ecology and Management, vol.274, pp.143-150, 2012.
DOI : 10.1016/j.foreco.2012.02.033

J. Camarero, C. Bigler, J. Linares, and E. Gil-pelegrín, Synergistic effects of past historical logging and drought on the decline of Pyrenean silver fir forests, Forest Ecology and Management, vol.262, issue.5, pp.759-769, 2011.
DOI : 10.1016/j.foreco.2011.05.009