A. Adamou, Mobilité résidentielle et processus d'e ´talement de la ville de Niamey (Niger), 2012.

N. Grimm, S. Faeth, N. Golubiewskin, C. Redman, and J. Wu, Global Change and the Ecology of Cities, Science, vol.319, issue.5864, pp.756-760, 2008.
DOI : 10.1126/science.1150195

M. Luniak, Synurbization -adaptation of animal wildlife to urban development, pp.50-55, 2004.

I. Fernandez and J. Simonetti, Small mammal assemblages in fragmented shrublands of urban areas of Central Chile, Urban Ecosystems, vol.30, issue.2, pp.377-387, 2013.
DOI : 10.1007/s11252-012-0272-1

D. Wilson and D. Reeder, Mammal species of the worl: a taxonomic and geographic reference 3rd edition, 2005.

F. Audoin-rouzeau and J. Vigne, La Colonisation de l'Europe par le rat noir (Rattus rattus), Rev Pal, vol.13, pp.125-145, 1994.

K. Aplin, H. Suzuki, A. Chinen, R. Chesser, and J. Ten-have, Multiple Geographic Origins of Commensalism and Complex Dispersal History of Black Rats, PLoS ONE, vol.55, issue.11, p.26357, 2011.
DOI : 10.1371/journal.pone.0026357.s007
URL : https://hal.archives-ouvertes.fr/ird-00714383

F. Bonhomme, A. Orth, T. Cucchi, H. Rajabi-maham, and J. Catalan, Genetic differentiation of the house mouse around the Mediterranean basin: matrilineal footprints of early and late colonization, Proceedings of the Royal Society B: Biological Sciences, vol.17, issue.24, pp.1034-1043, 2011.
DOI : 10.1093/molbev/msj096

E. Matisoo-smith and J. Robins, From The Cover: Origins and dispersals of Pacific peoples: Evidence from mtDNA phylogenies of the Pacific rat, Proceedings of the National Academy of Sciences, vol.101, issue.24, pp.9167-9172, 2004.
DOI : 10.1073/pnas.0403120101

A. Bastos, D. Nair, P. Taylor, H. Brettschneider, and F. Kirsten, Genetic monitoring detects an overlooked cryptic species and reveals the diversity and distribution of three invasive Rattus congeners in south Africa, BMC Genetics, vol.12, issue.1, p.26, 2011.
DOI : 10.1007/BF02602930

J. Lack, M. Hamilton, J. Braun, M. Mares, and R. Van-den-busche, Comparative phylogeography of invasive Rattus rattus and Rattus norvegicus in the U.S. reveals distinct colonization histories and dispersal, Biological Invasions, vol.26, issue.5, pp.1067-1087, 2013.
DOI : 10.1007/s10530-012-0351-5

C. Himsworth, K. Parsons, C. Jardine, and D. Patrick, Rats, Cities, People, and Pathogens: A Systematic Review and Narrative Synthesis of Literature Regarding the Ecology of Rat-Associated Zoonoses in Urban Centers, Vector-Borne and Zoonotic Diseases, vol.13, issue.6, pp.349-359, 2013.
DOI : 10.1089/vbz.2012.1195

G. Meerburg, G. Singleton, and A. Kijlstra, Rodent-borne diseases and their risks for public health, Critical Reviews in Microbiology, vol.24, issue.4, pp.221-270, 2009.
DOI : 10.1080/01652176.1987.9694138

S. Langton, D. Cowan, and A. Meyer, The occurrence of commensal rodents in dwellings as revealed by the 1996 English House Condition Survey, Journal of Applied Ecology, vol.38, issue.4, pp.699-709, 2001.
DOI : 10.1046/j.1365-2664.2001.00631.x

G. Murphy, P. Marshall, G. Singleton, L. Hinds, C. Krebs et al., Factors influencing the occurrence of rodent infestations in an inner city area of Manchester Rats mice and people: rodent biology and management, Australian Center for International Agricultural Research Canberra, pp.469-472, 2003.

P. Promkerd, Y. Khoprasert, P. Virathavone, M. Thoummabouth, and O. Sirisak, Factors explaining the abundance of rodents in the city of Luang Prabang, Lao PDR, as revealed by field and household surveys, Integrative Zoology, vol.40, issue.1, pp.11-20, 2008.
DOI : 10.1111/j.1749-4877.2008.00069.x

P. Taylor, L. Arntzen, M. Hayter, M. Iles, and J. Frean, Understanding and managing sanitary risks due to rodent zoonoses in an African city: beyond the Boston Model, Integrative Zoology, vol.24, issue.Suppl., pp.38-50, 2008.
DOI : 10.1111/j.1749-4877.2008.00072.x

R. Cavia, G. Cueto, and O. Suarez, Changes in rodent communities according to the landscape structure in an urban ecosystem, Landscape and Urban Planning, vol.90, issue.1-2, pp.11-19, 2009.
DOI : 10.1016/j.landurbplan.2008.10.017

G. Houéménou, A. Ahmed, R. Libois, and R. Hartskeerl, Leptospira spp. prevalence in small mammal populations in Cotonou, Benin. ISRN Epidemiol, 2013.

C. Himsworth, J. Bidulka, K. Parsons, A. Feng, and P. Tang, Ecology of Leptospira interrogans in Norway Rats (Rattus norvegicus) in an Inner-City Neighborhood of Vancouver, Canada, PLoS Neglected Tropical Diseases, vol.2, issue.6, p.2270, 2013.
DOI : 10.1371/journal.pntd.0002270.s001

A. Feng and C. Himsworth, The secret life of the city rat: a review of the ecology of urban Norway and black rats (Rattus norvegicus and Rattus rattus), Urban Ecosystems, vol.118, issue.1, 2013.
DOI : 10.1007/s11252-013-0305-4

M. Gomez, M. Provensal, and J. Polop, Effect of interspecific competition on Mus musculus in an urban area, Journal of Pest Science, vol.135, issue.4, pp.235-240, 2008.
DOI : 10.1007/s10340-008-0212-2

A. Salifou, Histoire du Niger: e ´poques précoloniale et coloniale, p.317, 2010.

H. Sidikou, Notes sur l9Histoire de Niamey, Niameyà360u.Niamey Niger, 2011.

M. Garba, Rongeurs urbains et invasion biologique dans le sud-ouest du Niger: e ´cologie des communautés et génétique des populations, p.261, 2012.

L. Granjon and J. Duplantier, Les rongeurs de l'Afrique sahélo-soudanienne, Publications Scientifiques IRD, p.215, 2009.

A. Olayemi, V. Nicolas, J. Hulselmans, D. Missoup, and E. Fichet-calvet, Taxonomy of the African giant pouched rats (Nesomyidae: Cricetomys): molecular and craniometric evidence support an unexpected high species diversity, Zoological Journal of the Linnean Society, vol.165, issue.3, pp.700-719, 2012.
DOI : 10.1111/j.1096-3642.2012.00823.x

F. Veyrunes, J. Britton-davidian, T. Robinson, E. Fichet-calvet, and C. Denys, Molecular phylogeny of the African pygmy mice, subgenus Nannomys (Rodentia, Murinae, Mus): Implications for chromosomal evolution, Molecular Phylogenetics and Evolution, vol.36, issue.2, pp.358-369, 2005.
DOI : 10.1016/j.ympev.2005.02.011

G. Dobigny, A. Nomao, and J. Gautun, A cytotaxonomic survey of Rodents from Niger: implications for systematics, biodiversity and biogeography, Mammalia, vol.66, issue.4, pp.495-523, 2002.
DOI : 10.1515/mamm.2002.66.4.495

G. Dobigny, E. Lecompte, C. Tatard, P. Gauthier, and K. Bâ, (Muridae, Murinae) using combined cytogenetic and molecular data, Journal of Zoology, vol.9, issue.(Suppl. 1), pp.368-374, 2008.
DOI : 10.1111/j.1469-7998.2008.00500.x
URL : https://hal.archives-ouvertes.fr/hal-00351317

E. Lecompte, C. Brouat, J. Duplantier, M. Galan, and L. Granjon, Molecular identification of four cryptic species of Mastomys (Rodentia, Murinae), Biochemical Systematics and Ecology, vol.33, issue.7, pp.681-689, 2005.
DOI : 10.1016/j.bse.2004.12.015

M. Galan, V. Hooft, W. Legrand, D. Berthier, K. Loiseau et al., A multiplex panel of microsatellite markers for widespread sub-Saharan rodents of the genus Mastomys, Molecular Ecology Notes, vol.127, issue.3, pp.321-323, 2004.
DOI : 10.1111/j.1471-8286.2004.00681.x

A. Gilabert, A. Loiseau, J. Duplantier, S. Rahelinira, and L. Rahalison, Genetic structure of black rat populations in a rural plague focus in Madagascar, Canadian Journal of Zoology, vol.85, issue.9, pp.965-972, 2007.
DOI : 10.1139/Z07-083
URL : https://hal.archives-ouvertes.fr/ird-00261411

A. Loiseau, S. Rahelinirina, L. Rahalison, A. Konecny, and J. Duplantier, Isolation and characterization of microsatellites in Rattus rattus, Mol Ecol Res, vol.8, pp.915-918, 2008.

N. Gotelli, NULL MODEL ANALYSIS OF SPECIES CO-OCCURRENCE PATTERNS, Ecology, vol.81, issue.9, pp.2606-2621, 2000.
DOI : 10.2307/1943025

N. Gotelli and G. Entsminger, EcoSim: Null models software for ecology. Version 7.72 Acquired Intelligence Inc. and Kesey-Bear, 2006.

L. Stone and A. Roberts, The checkerboard score and species distributions, Oecologia, vol.11, issue.1, pp.74-79, 1990.
DOI : 10.1007/BF00317345

D. Mackenzie, J. Nichols, G. Lachman, S. Droege, A. Royle et al., ESTIMATING SITE OCCUPANCY RATES WHEN DETECTION PROBABILITIES ARE LESS THAN ONE, Ecology, vol.83, issue.8, pp.2248-2255, 2002.
DOI : 10.1016/S0169-5347(01)02205-4

D. Mackenzie, Occupancy estimation and modeling: inferring patterns and dynamics of species occurrence, p.324, 2006.

P. Goovaerts, Geostatistics for natural resources evaluation, 1997.

A. Journel and C. Huijbregts, Mining geostatistics, 1978.

G. Robertson, Geostatistics in Ecology: Interpolating With Known Variance, Ecology, vol.68, issue.3, pp.744-748, 1987.
DOI : 10.2307/1938482

E. Isaaks and R. Srivastava, Applied geostatistics, p.561, 1989.

P. Legendre and L. Legendre, Numerical Ecology, 1998.
URL : https://hal.archives-ouvertes.fr/hal-00530195

R. Webster and M. Oliver, Statistical Methods in Soil and Land Resource Survey., Biometrics, vol.49, issue.4, p.316, 1990.
DOI : 10.2307/2532283

M. Fortin and M. Dale, Spatial analysis: a guide for ecologists, p.365, 2005.
DOI : 10.1017/CBO9780511542039

P. Goovaerts, Geostatistics in soil science: state-of-the-art and perspectives, Geoderma, vol.89, issue.1-2, pp.1-45, 1999.
DOI : 10.1016/S0016-7061(98)00078-0

P. Goovaerts, Geostatistical tools for characterizing the spatial variability of microbiological and physico-chemical soil properties, Biology and Fertility of Soils, vol.27, issue.4, pp.315-334, 1998.
DOI : 10.1007/s003740050439

G. Houéménou, Les petits mammifères de la ville de Cotonou (Bénin) pourraient-ils constituer un risque pour la santé humaine?, p.pp, 2013.

A. Bradman, J. Chevrier, I. Tager, M. Lipsett, and J. Sedgwick, Association of Housing Disrepair Indicators with Cockroach and Rodent Infestations in a Cohort of Pregnant Latina Women and Their Children, Environmental Health Perspectives, vol.113, issue.12, pp.1795-1801, 2005.
DOI : 10.1289/ehp.7588

J. Panti-may, S. Hernandez-betancourt, H. Ruiz-pina, and S. Medina-peralta, Abundance and population parameters of commensal rodents present in rural households in Yucatan, Mexico, International Biodeterioration & Biodegradation, vol.66, issue.1, pp.77-81, 2012.
DOI : 10.1016/j.ibiod.2011.10.006

E. Omudu and T. Ati, A Survey of rats trapped in residential apartments and their ectoparasites in Makurdi Nigeria, Res J Agric Biol Sci, vol.6, pp.144-149, 2010.

S. Lavergne and J. Molofsky, Increased genetic variation and evolutionary potential drive the success of an invasive grass, Proceedings of the National Academy of Sciences, vol.104, issue.10, pp.3883-3888, 2007.
DOI : 10.1073/pnas.0607324104
URL : https://hal.archives-ouvertes.fr/halsde-00293172

C. Kolar and D. Lodge, Progress in invasion biology: predicting invaders, Trends in Ecology & Evolution, vol.16, issue.4, pp.199-204, 2001.
DOI : 10.1016/S0169-5347(01)02101-2

J. Duplantier, L. Granjon, F. Adam, and K. Bâ, Répartition actuelle du rat noir (Rattus rattus) au Sénégal: Facteurs historiques et e ´cologiques, pp.339-346, 1991.

J. Duplantier, L. Granjon, and K. Bâ, Répartition biogéographique des petits rongeurs au Sénégal, J Afr Zool, vol.111, pp.17-26, 1997.

P. Kalémé, J. Bates, H. Balesi, M. Gambalemoke, and J. Kerbis-peterhans, Origin and putative colonisation routes for invasive rodent taxa in Democratic Republic of Congo, Afr Zool, vol.40, pp.133-145, 2011.

A. Konecny, A. Estoup, J. Duplantier, J. Bryja, and K. Bâ, ) in Senegal, West Africa, Molecular Ecology, vol.24, issue.83, pp.286-300, 2013.
DOI : 10.1111/mec.12112

S. Belmain, A. Meyer, R. Timbrine, L. Pinecela, G. Singleton et al., Managing rodent pests in households and food stores through intensive trapping) Rats, mice and people: rodent biology and management, Australian Center for International Agricultural Research, pp.440-445, 2003.

A. Monadjem, T. Mahlaba, N. Dlamini, S. Eiseb, and S. Belmain, Impact of crop cycle on movement patterns of pest rodent species between fields and houses in Africa, Wildlife Research, vol.38, issue.7, pp.603-609, 2011.
DOI : 10.1071/WR10130

L. Granjon and G. Cheylan, Le sort de rats noirs (Rattus rattus) introduits sur une ? ?le, révélé par radio-tracking, C R Acad Sci Paris, vol.309, pp.571-575, 1989.

M. Garba and G. Dobigny, Abstract, Mammalia, vol.78, issue.2, 2014.
DOI : 10.1515/mammalia-2013-0042

P. Daszak, A. Cunningham, and A. Hyatt, Emerging Infectious Diseases of Wildlife-- Threats to Biodiversity and Human Health, Science, vol.287, issue.5452, pp.443-449, 2000.
DOI : 10.1126/science.287.5452.443

J. Pickering and C. Norris, New evidence concerning the extinction of the endemic Murid Rattus macleari from Christmas island indian ocean, Austral Mammal, vol.19, pp.19-25, 1996.

K. Wyatt, P. Campos, M. Gilbert, S. Kolokotronis, and W. Hynes, Historical Mammal Extinction on Christmas Island (Indian Ocean) Correlates with Introduced Infectious Disease, PLoS ONE, vol.10, issue.11, p.3602, 2008.
DOI : 10.1371/journal.pone.0003602.s005

G. Dobigny, P. Poirier, K. Hima, O. Cabaret, and P. Gauthier, Molecular survey of rodent-borne Trypanosoma in Niger with special emphasis on T. lewisi imported by invasive black rats, Acta Tropica, vol.117, issue.3, pp.183-188, 2011.
DOI : 10.1016/j.actatropica.2010.11.004
URL : https://hal.archives-ouvertes.fr/hal-01113935

M. Garba, M. Kane, S. Gagaré, I. Kadaouré, and R. Sidikou, Local perception of rodent-associated problems in Sahelian urban areas: a survey in Niamey, Niger, Urban Ecosystems, vol.29, issue.2, 2014.
DOI : 10.1007/s11252-013-0336-x

D. Djidingar, J. Chippaux, G. Gragnic, O. Tchani, D. Meynard et al., La leishmaniose viscérale au Niger: six nouveaux cas confirmés parasitologiquement, Bull Soc Path Ex, vol.90, pp.27-29, 1997.

A. Mercier, M. Garba, H. Bonnabau, M. Kane, and J. Rossi, Toxoplasmosis seroprevalence in urban rodents: a survey in Niamey, Niger, Mem??rias do Instituto Oswaldo Cruz, vol.108, issue.4, pp.399-407, 2013.
DOI : 10.1590/S0074-0276108042013002
URL : https://hal.archives-ouvertes.fr/hal-01219025

S. Howie, M. Guy, L. Fleming, W. Bailey, and H. Noyes, A Gambian Infant with Fever and an Unexpected Blood Film, PLoS Medicine, vol.410, issue.9, p.3553, 2006.
DOI : 10.1371/journal.pmed.0030355.g004

W. Bergmans, Les rongeurs du Bénin: espèces trouvées et espèces attendues, pp.23-44, 1999.

F. Angelici and L. Luiselli, Patterns of specific diversity and population size in small mammals from arboreal and ground-dwelling guilds of a forest area in southern Nigeria, Journal of Zoology, vol.143, issue.1, pp.9-16, 2005.
DOI : 10.1086/282398

I. Olaseha, M. Sridhar, P. Obiako, and A. Oladapp, Rat Infestations in Urban and Rural Areas in Nigeria: public health implications, The Journal of the Royal Society for the Promotion of Health, vol.114, issue.6, pp.300-303, 1994.
DOI : 10.1177/146642409411400604

D. Rosevear, The rodents of West Africa, British Museum of Natural History, p.604, 1969.