S. Aaronson, T. Berner, K. Gold, L. Kushner, N. J. Patni et al., and its bloom form, Journal of Plankton Research, vol.5, issue.5, pp.693-700, 1983.
DOI : 10.1093/plankt/5.5.693

P. G. Appleby and F. Oldfield, The calculation of lead-210 dates assuming a constant rate of supply of unsupported 210Pb to the sediment, CATENA, vol.5, issue.1, pp.1-8, 1978.
DOI : 10.1016/S0341-8162(78)80002-2

K. Ashok, S. K. Behera, S. A. Rao, H. Weng, and T. Yamagata, El Ni??o Modoki and its possible teleconnection, Journal of Geophysical Research, vol.10, issue.D8, 2007.
DOI : 10.1029/2006JC003798

A. R. Atwood and J. P. Sachs, Separating ITCZ- and ENSO-related rainfall changes in the Gal??pagos over the last 3 kyr using D/H ratios of multiple lipid biomarkers, Earth and Planetary Science Letters, vol.404, pp.408-419, 2014.
DOI : 10.1016/j.epsl.2014.07.038

J. H. Belcher, Notes on the physiology of Botryococcus braunii Kützing, Arch. Microbiol, vol.61, pp.335-346, 1968.

P. Braconnot, Y. Luan, S. Brewer, and W. Zheng, Impact of Earth???s orbit and freshwater fluxes on Holocene climate mean seasonal cycle and ENSO characteristics, Climate Dynamics, vol.30, issue.7???8, pp.1081-1092, 2012.
DOI : 10.1007/s00382-011-1029-x

C. Bronk-ramsey, Deposition models for chronological records, Quaternary Science Reviews, vol.27, issue.1-2, pp.42-60, 2008.
DOI : 10.1016/j.quascirev.2007.01.019

A. C. Clement, R. Seager, and M. A. Cane, Orbital controls on the El Ni??o/Southern Oscillation and the tropical climate, Paleoceanography, vol.10, issue.4, pp.441-456, 1999.
DOI : 10.1029/1999PA900013

A. C. Clement, R. Seager, and M. A. Cane, Suppression of El Ni??o during the Mid-Holocene by changes in the Earth's orbit, Paleoceanography, vol.115, issue.6, pp.731-737, 2000.
DOI : 10.1029/1999PA000466

K. M. Cobb, C. D. Charles, H. Cheng, and R. L. Edwards, El Ni??o/Southern Oscillation and tropical Pacific climate during the last millennium, Nature, vol.424, issue.6946, pp.271-276, 2003.
DOI : 10.1038/nature01779

K. M. Cobb, N. Westphal, H. R. Sayani, J. T. Watson, E. D. Lorenzo et al., Highly Variable El Nino-Southern Oscillation Throughout the Holocene, Highly variable El Niño-Southern Oscillation throughout the Holocene, pp.67-70, 2013.
DOI : 10.1126/science.1228246

P. A. Colinvaux, Climate and the Galapagos Islands, Nature, vol.125, issue.5375, pp.17-20, 1972.
DOI : 10.1038/240017a0

M. Collins, The impact of global warming on the tropical Pacific Ocean and El Ni??o, Nature Geoscience, vol.22, issue.6, pp.391-397, 2010.
DOI : 10.1038/ngeo868

J. L. Conroy, J. T. Overpeck, J. E. Cole, T. M. Shanahan, and M. Steinitz-kannan, Holocene changes in eastern tropical Pacific climate inferred from a Gal??pagos lake sediment record, Quaternary Science Reviews, vol.27, issue.11-12, pp.1166-1180, 2008.
DOI : 10.1016/j.quascirev.2008.02.015

W. Dansgaard, Stable isotopes in precipitation, Tellus, vol.16, pp.436-468, 1964.

P. N. Dinezio, A. C. Clement, G. A. Vecchi, B. J. Soden, B. P. Kirtman et al., Climate Response of the Equatorial Pacific to Global Warming, Journal of Climate, vol.22, issue.18, pp.4873-4892, 2009.
DOI : 10.1175/2009JCLI2982.1

J. Emile-geay, M. Cane, R. Seager, A. Kaplan, and P. Almasi, El Ni??o as a mediator of the solar influence on climate, Paleoceanography, vol.18, issue.12, 2007.
DOI : 10.1029/2006PA001304

J. Emile-geay, K. M. Cobb, M. E. Mann, and A. T. Wittenberg, Estimating Central Equatorial Pacific SST Variability over the Past Millennium. Part II: Reconstructions and Implications, Journal of Climate, vol.26, issue.7, pp.2329-2352, 2013.
DOI : 10.1175/JCLI-D-11-00511.1

J. Green, S. A. Corbet, and E. Betney, Ecological studies on crater lakes in West Cameroon. Debundsha Lake, Journal of Zoology, vol.13, issue.1, pp.199-223, 1974.
DOI : 10.1111/j.1469-7998.1974.tb03126.x

G. J. Gu, R. F. Adler, and A. H. Sobel, The Eastern Pacific ITCZ during the Boreal Spring, Journal of the Atmospheric Sciences, vol.62, issue.4, pp.1157-1174, 2005.
DOI : 10.1175/JAS3402.1

J. D. Haigh, The Impact of Solar Variability on Climate, Science, vol.272, issue.5264, pp.981-984, 1996.
DOI : 10.1126/science.272.5264.981

L. L. Hood, Solar variability and its effect on the Earth's atmospheric and climate system, AGU, pp.283-303, 2004.

L. L. Hood and B. E. Soukharev, The Lower-Stratospheric Response to 11-Yr Solar Forcing: Coupling to the Troposphere???Ocean Response, Journal of the Atmospheric Sciences, vol.69, issue.6, pp.1841-1864, 2012.
DOI : 10.1175/JAS-D-11-086.1

E. Kebede and A. Belay, Species composition and phytoplankton biomass in a tropical African lake (Lake Awassa, Ethiopia), Hydrobiologia, vol.3, issue.2, pp.13-32, 1994.
DOI : 10.1007/BF00006802

W. S. Kessler, The circulation of the eastern tropical Pacific: A review, Progress in Oceanography, vol.69, issue.2-4, pp.181-217, 2006.
DOI : 10.1016/j.pocean.2006.03.009

A. Koutavas and S. Joanides, El Ni??o-Southern Oscillation extrema in the Holocene and Last Glacial Maximum, Paleoceanography, vol.30, issue.1, 2012.
DOI : 10.1007/s00382-007-0320-3

N. A. Krivova and S. K. Solanki, Reconstruction of solar UV irradiance, Advances in Space Research, vol.35, issue.3, pp.361-364, 2005.
DOI : 10.1016/j.asr.2004.12.027

N. Kurita, K. Ichiyanagi, J. Matsumoto, M. D. Yamanaka, and T. Ohata, The relationship between the isotopic content of precipitation and the precipitation amount in tropical regions, Journal of Geochemical Exploration, vol.102, issue.3, pp.113-122, 2009.
DOI : 10.1016/j.gexplo.2009.03.002

M. Latif and A. Grötzner, The equatorial Atlantic oscillation and its response to ENSO, Climate Dynamics, vol.16, issue.2-3, pp.213-218, 2000.
DOI : 10.1007/s003820050014

G. Leduc, L. Vidal, O. Cartapanis, and E. Bard, Modes of eastern equatorial Pacific thermocline variability: Implications for ENSO dynamics over the last glacial period, Paleoceanography, vol.30, issue.1, 2009.
DOI : 10.1007/s00382-007-0320-3

G. Leduc, L. Vidal, K. Tachikawa, and E. Bard, Abstract, Quaternary Research, vol.173, issue.01, pp.123-131, 2009.
DOI : 10.1038/364218a0

W. T. Liu and X. Xie, Double intertropical convergence zones-a new look using scatterometer, Geophysical Research Letters, vol.24, issue.11, 2002.
DOI : 10.1029/2002GL015431

T. M. Marchitto, R. Muscheler, J. D. Ortiz, J. D. Carriquiry, and A. Van-geen, Dynamical Response of the Tropical Pacific Ocean to Solar Forcing During the Early Holocene, Science, vol.330, issue.6009, pp.1378-1381, 2010.
DOI : 10.1126/science.1194887

F. G. Mccormac, Radiocarbon Calibration from 0?26 cal kyr BP? SHCa104 Southern Hemisphere Calibration, pp.0-11, 2004.

H. V. Mcgregor, M. J. Fischer, M. K. Gagan, D. Fink, S. J. Phipps et al., A weak El Ni??o/Southern Oscillation with delayed seasonal growth around 4,300 years ago, Nature Geoscience, vol.21, issue.11, pp.949-953, 2013.
DOI : 10.1038/ngeo1936

G. A. Meehl and J. M. Arblaster, A Lagged Warm Event???Like Response to Peaks in Solar Forcing in the Pacific Region, Journal of Climate, vol.22, issue.13, pp.3647-3660, 2009.
DOI : 10.1175/2009JCLI2619.1

G. A. Meehl, J. M. Arblaster, K. Matthes, F. Sassi, and H. Van-loon, Amplifying the Pacific Climate System Response to a Small 11-Year Solar Cycle Forcing, Science, vol.325, issue.5944, pp.1114-1118, 2009.
DOI : 10.1126/science.1172872

P. Metzger, E. Casadevall, M. J. Pouet, and Y. Pouet, Structures of some botryococcenes: branched hydrocarbons from the b-race of the green alga Botryococcus braunii, Phytochemistry, vol.24, issue.12, pp.2995-3002, 1985.
DOI : 10.1016/0031-9422(85)80043-1

X. Miao, J. A. Mason, J. B. Swinehart, D. B. Loope, P. R. Hanson et al., A 10,000 year record of dune activity, dust storms, and severe drought in the central Great Plains, Geology, vol.35, issue.2, pp.119-122, 2007.
DOI : 10.1130/G23133A.1

C. M. Moy, G. O. Seltzer, D. T. Rodbell, and D. M. Anderson, Variability of El Ni??o/Southern Oscillation activity at millennial timescales during the Holocene epoch, Nature, vol.35, issue.6912, pp.162-165, 2002.
DOI : 10.1006/qres.1993.1040

M. Pascolini-campbell, D. Zanchettin, O. Bothe, C. Timmreck, D. Matei et al., Toward a record of Central Pacific El Ni??o events since 1880, Theoretical and Applied Climatology, vol.10, issue.7263, pp.704-718, 2014.
DOI : 10.1007/s00704-014-1114-2

Z. Zhang, El Ni??o evolution during the Holocene revealed by a biomarker rain gauge in the Gal??pagos Islands, Earth and Planetary Science Letters, vol.404, pp.420-434, 2014.
DOI : 10.1016/j.epsl.2014.07.013

L. C. Peterson, G. H. Haug, K. A. Hughen, and U. Rohl, Rapid Changes in the Hydrologic Cycle of the Tropical Atlantic During the Last Glacial, Science, vol.290, issue.5498, pp.1947-1951, 2000.
DOI : 10.1126/science.290.5498.1947

R. T. Pierrehumbert, Climate change and the tropical Pacific: The sleeping dragon wakes, Proc. Natl. Acad. Sci. USA 97, pp.1355-1358, 2000.
DOI : 10.1073/pnas.97.4.1355

. Pryet, C. Domínguez, A. Tomai, P. F. Chaumont, C. Ozouville et al., Quantification of cloud water interception along the windward slope of Santa Cruz Island, Galapagos (Ecuador), Agricultural and Forest Meteorology, vol.161, pp.94-106, 2012.
DOI : 10.1016/j.agrformet.2012.03.018

URL : https://hal.archives-ouvertes.fr/hal-00694753

E. M. Rasmusson and T. H. Carpenter, Variations in Tropical Sea Surface Temperature and Surface Wind Fields Associated with the Southern Oscillation/El Ni??o, Monthly Weather Review, vol.110, issue.5, pp.354-384, 1982.
DOI : 10.1175/1520-0493(1982)110<0354:VITSST>2.0.CO;2

P. J. Richerson, P. J. Neale, W. Wurtsbaugh, R. Alfaro, and W. Vincent, Patterns of temporal variation in Lake Titicaca. A high altitude tropical lake. I. Background, physical and chemical processes, and primary production, Hydrobiologia, vol.19, issue.1, pp.205-220, 1986.
DOI : 10.1007/BF00027241

C. Risi, S. Bony, and F. Vimeux, Influence of convective processes on the isotopic composition (? 18 O and ?D) of precipitation and water vapor in the tropics: 2. Physical interpretation of the amount effect, J. Geophys. Res, vol.113, 2008.

J. P. Sachs, D. Sachse, R. H. Smittenberg, Z. H. Zhang, D. S. Battisti et al., Southward movement of the Pacific intertropical convergence zone AD???1400???1850, Nature Geoscience, vol.42, issue.8, pp.519-525, 2009.
DOI : 10.1038/ngeo554

O. R. Salau, B. Schneider, W. Park, V. Khon, and M. Latif, Modeling the ENSO impact of orbitally induced mean state climate changes, Journal of Geophysical Research: Oceans, vol.30, issue.10, 2012.
DOI : 10.1007/s00382-007-0320-3

D. H. Sandweiss, J. B. Richardson, E. J. Reitz, H. B. Rollins, and K. A. Maasch, Geoarchaeological Evidence from Peru for a 5000 Years B.P. Onset of El Nino, Science, vol.273, issue.5281, pp.1531-1533, 1996.
DOI : 10.1126/science.273.5281.1531

M. Steinitz-kannan, M. A. Riedinger, W. Last, M. Brenner, and M. C. Miller, A 6000-year record of intense evidence of El Niño from sediments in a Galápagos Islands lake, Bull. Inst. Fr. Études Andines, vol.27, pp.581-592, 1998.

G. T. Taylor, Ecosystem responses in the southern Caribbean Sea to global climate change, Proc. Natl. Acad. Sci. USA, 2012.
DOI : 10.1073/pnas.1207514109

K. Thirumalai, J. W. Partin, C. S. Jackson, and T. M. Quinn, Statistical constraints on El Ni??o Southern Oscillation reconstructions using individual foraminifera: A sensitivity analysis, Paleoceanography, vol.365, issue.1856, 2013.
DOI : 10.1098/rsta.2007.2045

S. A. Townsend, Perennial domination of phytoplankton by Botryococcus and Peridinium in a discontinuously polymictic reservoir (tropical Australia), Arch. Hydrobiol, vol.151, pp.529-548, 2001.

K. E. Trenberth and C. J. Guillemot, Physical Processes Involved in the 1988 Drought and 1993 Floods in North America, Journal of Climate, vol.9, issue.6, pp.1288-1298, 1996.
DOI : 10.1175/1520-0442(1996)009<1288:PPIITD>2.0.CO;2

M. Trueman and N. Ozouville, Characterizing the Galápagos terrestrial climate in the face of global climate change, Galápagos Res, vol.67, pp.26-37, 2010.

H. Van-loon and G. A. Meehl, The average influence of decadal solar forcing on the atmosphere in the South Pacific region, Geophysical Research Letters, vol.23, issue.4, 2011.
DOI : 10.1029/2011GL047794

M. Vonmoos, J. Beer, and R. Muscheler, Be in the Greenland Ice Core Project ice core, Journal of Geophysical Research, vol.102, issue.A9, 2006.
DOI : 10.1029/2005JA011500

L. V. Wake and L. W. Hillen, Study of a ?bloom? of the oil-rich algaBotryococcus braunii in the Darwin River Reservoir, Biotechnology and Bioengineering, vol.3, issue.8, pp.1637-1656, 1980.
DOI : 10.1002/bit.260220808

Z. H. Zhang and J. P. Sachs, Hydrogen isotope fractionation in freshwater algae: I. Variations among lipids and species, Organic Geochemistry, vol.38, issue.4, pp.582-608, 2007.
DOI : 10.1016/j.orggeochem.2006.12.004

Z. H. Zhang, P. Metzger, and J. P. Sachs, Biomarker evidence for the co-occurrence of three races (A, B and L) of Botryococcus braunii in El Junco Lake, Gal??pagos, Organic Geochemistry, vol.38, issue.9, pp.1459-1478, 2007.
DOI : 10.1016/j.orggeochem.2007.05.015

Z. Zhang, J. P. Sachs, and A. Marchetti, Hydrogen isotope fractionation in freshwater and marine algae: II. Temperature and nitrogen limited growth rate effects, Organic Geochemistry, vol.40, issue.3, pp.428-439, 2009.
DOI : 10.1016/j.orggeochem.2008.11.002

Z. H. Zhang, P. Metzger, and J. P. Sachs, Co-occurrence of long chain diols, keto-ols, hydroxy acids and keto acids in recent sediments of Lake El Junco, Gal??pagos Islands, Organic Geochemistry, vol.42, issue.7, pp.823-837, 2011.
DOI : 10.1016/j.orggeochem.2011.04.012

Z. H. Zhang, P. Metzger, and J. P. Sachs, Unprecedented long chain 1-chloroalkenes and 1-chloroalkanes in the Holocene sediments of Lake El Junco, Gal??pagos Islands, Organic Geochemistry, vol.57, pp.1-6, 2013.
DOI : 10.1016/j.orggeochem.2013.01.009

Z. H. Zhang, P. Metzger, and J. P. Sachs, Bound lipid biomarkers in sediments from El Junco Lake, Gal??pagos Islands, Organic Geochemistry, vol.75, pp.122-128, 2014.
DOI : 10.1016/j.orggeochem.2014.06.013

W. Zheng, P. Braconnot, E. Guilyardi, U. Merkel, and Y. Yu, ENSO at 6ka and 21ka from ocean???atmosphere coupled model simulations, Climate Dynamics, vol.28, issue.23, pp.745-762, 2008.
DOI : 10.1007/s00382-007-0320-3

URL : https://hal.archives-ouvertes.fr/hal-00770613