B. Amati and S. M. Gasser, Chromosomal ARS and CEN elements bind specifically to the yeast nuclear scaffold, Cell, vol.54, issue.7, pp.967-978, 1988.
DOI : 10.1016/0092-8674(88)90111-0

B. Amati, L. Pick, T. Laroche, and S. M. Gasser, Nuclear scaffold attachment stimulates, but is not essential for ARS activity in Saccharomyces cerevisiae: analysis of the Drosophila ftz SAR, EMBO J, vol.9, pp.4007-4016, 1990.

G. Barth and C. Gaillardin, Yarrowia lipolytica, Nonconventional yeasts in biotechnology?a handbook, pp.313-388, 1996.

M. Bénard, C. Lagnel, D. Pallotta, and G. Pierron, Mapping of a replication origin within the promoter region of two unlinked, abundantly transcribed actin genes of Physarum polycephalum., Molecular and Cellular Biology, vol.16, issue.3, pp.968-976, 1996.
DOI : 10.1128/MCB.16.3.968

M. Bénard, C. Lagnel, and G. Pierron, rDNA, Nucleic Acids Research, vol.23, issue.9, pp.1447-1453, 1995.
DOI : 10.1093/nar/23.9.1447

A. I. Bogdanova, M. O. Agaphonov, and M. D. Ter-avanesyan, Plasmid reorganization during integrative transformation inHansenula polymorpha, Yeast, vol.8, issue.4, pp.343-353, 1995.
DOI : 10.1101/SQB.1983.047.01.132

B. J. Brewer and W. L. Fangman, Initiation preference at a yeast origin of replication., Proc. Natl. Acad. Sci. USA 91, pp.3418-3422, 1994.
DOI : 10.1073/pnas.91.8.3418

B. J. Brewer and W. L. Fangman, The localization of replication origins on ARS plasmids in S. cerevisiae, Cell, vol.51, issue.3, pp.463-471, 1987.
DOI : 10.1016/0092-8674(87)90642-8

C. Brun, Q. Dang, and R. Miassod, Studies of an 800-kilobase DNA stretch of the Drosophila X chromosome: comapping of a subclass of scaffold-attached regions with sequences able to replicate autonomously in Saccharomyces cerevisiae., Molecular and Cellular Biology, vol.10, issue.10, pp.5455-5463, 1990.
DOI : 10.1128/MCB.10.10.5455

URL : https://hal.archives-ouvertes.fr/hal-01596337

C. Brun, D. D. Dubey, and J. A. Huberman, pDblet, a stable autonomously replicating shuttle vector for Schizosaccharomyces pombe, Gene, vol.164, issue.1, pp.173-177, 1995.
DOI : 10.1016/0378-1119(95)00497-T

URL : https://hal.archives-ouvertes.fr/hal-01596234

C. Brun, N. Jullien, M. Jacobzone, C. Gautier, and R. Miassod, SARs, ARSs, and A,T-Rich Regions Evidenced by Restriction Mapping on an 835-kb DNA Fragment from Drosophila, Experimental Cell Research, vol.220, issue.2, pp.338-347, 1995.
DOI : 10.1006/excr.1995.1324

URL : https://hal.archives-ouvertes.fr/hal-01596335

W. C. Burhans and J. A. Huberman, DNA replication origins in animal cells: a question of context?, Science, vol.263, issue.5147, pp.639-640, 1994.
DOI : 10.1126/science.8303270

M. S. Caddle and M. P. Calos, Specific initiation at an origin of replication from Schizosaccharomyces pombe., Molecular and Cellular Biology, vol.14, issue.3, pp.1796-1805, 1994.
DOI : 10.1128/MCB.14.3.1796

R. K. Clyne and T. J. Kelly, Genetic analysis of an ARS element from the fission yeast Schizosaccharomyces pombe, EMBO J, vol.14, pp.6348-6357, 1995.

M. L. Depamphilis, Origins of DNA replication in metazoan chromosomes, J. Biol. Chem, vol.268, pp.1-4, 1993.

A. Dershowitz and C. S. Newlon, The effect on chromosome stability of deleting replication origins., Molecular and Cellular Biology, vol.13, issue.1, pp.391-398, 1993.
DOI : 10.1128/MCB.13.1.391

A. M. Deshpande and C. S. Newlon, The ARS consensus sequence is required for chromosomal origin function in Saccharomyces cerevisiae., Molecular and Cellular Biology, vol.12, issue.10, pp.4305-4313, 1992.
DOI : 10.1128/MCB.12.10.4305

J. F. Diffley, The initiation of DNA replication in the budding yeast cell division cycle, Yeast, vol.12, issue.16, pp.1651-1670, 1995.
DOI : 10.1128/MCB.12.10.4733

P. A. Dijkwel, J. P. Vaughn, and J. L. Hamlin, Replication initiation sites are distributed widely in the amplified CHO dihydrofolate reductase domain, Nucleic Acids Research, vol.22, issue.23, pp.4989-4996, 1994.
DOI : 10.1093/nar/22.23.4989

D. D. Dubey, L. R. Davis, S. A. Greenfeder, L. Y. Ong, J. Zhu et al., Evidence suggesting that the ARS elements associated with silencers of the yeast mating-type locus HML do not function as chromosomal DNA replication origins., Molecular and Cellular Biology, vol.11, issue.10, pp.5346-5355, 1991.
DOI : 10.1128/MCB.11.10.5346

D. D. Dubey, S. Kim, I. T. Todorov, and J. A. Huberman, Large, complex modular structure of a fission yeast DNA replication origin, Current Biology, vol.6, issue.4, pp.467-473, 1996.
DOI : 10.1016/S0960-9822(02)00514-6

D. D. Dubey, J. G. Zhu, D. L. Carlson, K. Sharma, and J. A. Huberman, Three ARS elements contribute to the ura4 replication origin region in the fission yeast, Schizosaccharomyces pombe, EMBO J, vol.13, pp.3638-3647, 1994.

L. Fabiani, M. Aragona, and L. Frontali, Isolation and sequence analysis of aK. lactis chromosomal DNA element able to autonomously replicate inS. cerevisiae andK. lactis, Yeast, vol.1, issue.1, pp.69-76, 1990.
DOI : 10.1016/0167-4781(89)90123-1

L. Fabiani, L. Frontali, and C. S. Newlon, Identification of an essential core element and stimulatory sequences in a Kluyveromyces lactis ARS element, KARS101, Molecular Microbiology, vol.19, issue.4, pp.757-766, 1996.
DOI : 10.1046/j.1365-2958.1996.401938.x

Q. Fan and M. Yao, New telomere formation coupled with site-specific chromosome breakage in Tetrahymena thermophila., Molecular and Cellular Biology, vol.16, issue.3, pp.1267-1274, 1996.
DOI : 10.1128/MCB.16.3.1267

URL : http://europepmc.org/articles/pmc231109?pdf=render

P. Fournier, L. Guyaneux, M. Chasles, and C. Gaillardin, Scarcity ofars sequences isolated in a morphogenesis mutant of the yeastYarrowia lipolytica, Yeast, vol.14, issue.1, pp.25-36, 1991.
DOI : 10.1271/bbb1961.48.1933

C. Gaillardin and A. Ribet, LEU2 directed expression of ?-galactosidase activity and phleomycin resistance in Yarrowia lipolytica, Current Genetics, vol.76, issue.5, pp.369-375, 1987.
DOI : 10.1007/BF00378179

A. E. Gammie and M. D. Rose, Identification and characterization of CEN12 in the budding yeast Saccharomyces cerevisiae, Current Genetics, vol.349, issue.6, pp.512-516, 1995.
DOI : 10.1007/BF00518162

G. P. Georgiev, Y. S. Vassetzky, A. N. Luchnik, V. V. Chernokhvostov, and S. V. Razin, Nuclear skeleton, DNA domains and control of replication and transcription, European Journal of Biochemistry, vol.9, issue.3, pp.613-624, 1991.
DOI : 10.1016/0006-291X(89)91978-5

D. M. Gilbert, H. Miyazawa, and M. L. Depamphilis, Site-specific initiation of DNA replication in Xenopus egg extract requires nuclear structure., Molecular and Cellular Biology, vol.15, issue.6, pp.2942-2954, 1995.
DOI : 10.1128/MCB.15.6.2942

S. A. Greenfeder and C. S. Newlon, Replication forks pause at yeast centromeres., Molecular and Cellular Biology, vol.12, issue.9, pp.4056-4066, 1992.
DOI : 10.1128/MCB.12.9.4056

URL : http://mcb.asm.org/content/12/9/4056.full.pdf

J. E. Haber and P. C. Thoburn, Healing of broken linear dicentric chromosomes in yeast, Genetics, vol.106, pp.207-226, 1984.

J. E. Haber, P. C. Thoburn, and D. Rogers, Meiotic and mitotic behavior of dicentric chromosomes in Saccharomyces cerevisiae, Genetics, vol.106, pp.185-205, 1984.

J. L. Hamlin, Mammalian origins of replication, BioEssays, vol.186, issue.10, pp.651-659, 1992.
DOI : 10.1128/MCB.8.12.5398

J. L. Hamlin and P. A. , On the nature of replication origins in higher eukaryotes, Current Opinion in Genetics & Development, vol.5, issue.2, pp.153-161, 1995.
DOI : 10.1016/0959-437X(95)80002-6

J. H. Hegemann and U. N. Fleig, The centromere of budding yeast, BioEssays, vol.9, issue.7, pp.451-460, 1993.
DOI : 10.1128/MCB.10.1.223

S. S. Heinzel, P. J. Krysan, C. T. Tran, and M. P. Calos, Autonomous DNA replication in human cells is affected by the size and the source of the DNA., Molecular and Cellular Biology, vol.11, issue.4, pp.2263-2272, 1991.
DOI : 10.1128/MCB.11.4.2263

J. J. Heus, B. J. Zonneveld, H. Y. De-steensma, J. A. Van-den, and . Berg, The consensus sequence of Kluyveromyces lactis centromeres show homology to functional centromeric DNA from Saccharomyces cerevisiae, Mol. Gen. Genet, vol.236, pp.355-362, 1993.

J. J. Heus, B. J. Zonneveld, H. Y. Steensma, and J. A. Vandenberg, Mutational analysis of centromeric DNA elements of Klayveromyces lactis and their role in determining the species specificity of the highly homologous centromeres from K. lactis and Saccharomyces cerevisiae, Molecular and General Genetics MGG, vol.6, issue.3, pp.325-333, 1994.
DOI : 10.1007/BF00301068

R. Huang and D. Kowalski, A DNA unwinding element and an ARS consensus comprise a replication origin within a yeast chromosome, EMBO J, vol.12, pp.4521-4531, 1993.

R. Huang and D. Kowalski, Multiple DNA Elements in ARS305 Determine Replication Origin Activity in a Yeast Chromosome, Nucleic Acids Research, vol.53, issue.3, pp.816-823, 1996.
DOI : 10.1016/0092-8674(88)90164-X

J. A. Huberman, Analysis of DNA replication origins and directions by two-dimensional gel electrophoresis The cell cycle: a practical approach, pp.213-234, 1994.

J. A. Huberman, L. D. Spotila, K. A. Nawotka, S. M. El-assouli, and L. R. Davis, The in vivo replication origin of the yeast 2??m plasmid, Cell, vol.51, issue.3, pp.473-481, 1987.
DOI : 10.1016/0092-8674(87)90643-X

O. Hyrien, C. Maric, and M. Méchali, Transition in Specification of Embryonic Metazoan DNA Replication Origins, Science, vol.270, issue.5238, pp.994-997, 1995.
DOI : 10.1126/science.270.5238.994

O. Hyrien and M. Méchali, eggs and egg extracts: a 2D gel electrophoretic analysis, Nucleic Acids Research, vol.20, issue.7, pp.1463-1469, 1992.
DOI : 10.1093/nar/20.7.1463

URL : https://academic.oup.com/nar/article-pdf/20/7/1463/4081735/20-7-1463.pdf

F. Iborra and M. M. Ball, Kluyveromyces marxianus small DNA fragments contain both autonomous replicative and centromeric elements that also function inKluyveromyces lactis, Yeast, vol.6, issue.12, pp.1621-1629, 1994.
DOI : 10.1128/MCB.10.8.3917

D. Jäger and P. Philippsen, Stabilization of dicentric chromosomes in Saccharomyces cerevisiae by telomere addition to broken ends or by centromere deletion, EMBO J, vol.8, pp.247-254, 1989.

F. Kirpekar, J. Friis, and K. Gullov, A search for an essential function of the replication originARS1 in the life cycle ofSaccharomyces cerevisiae, Yeast, vol.101, issue.4, pp.491-496, 1994.
DOI : 10.1128/MCB.12.9.4056

K. M. Kramer and J. E. Haber, New telomeres in yeast are initiated with a highly selected subset of TG1-3 repeats., Genes & Development, vol.7, issue.12a, pp.2345-2356, 1993.
DOI : 10.1101/gad.7.12a.2345

P. J. Krysan, J. G. Smith, and M. P. Calos, Autonomous replication in human cells of multimers of specific human and bacterial DNA sequences., Molecular and Cellular Biology, vol.13, issue.5, pp.2688-2696, 1993.
DOI : 10.1128/MCB.13.5.2688

M. H. Linskens and J. A. Huberman, Ambiguities in results obtained with 2D gel replicon mapping techniques, Nucleic Acids Research, vol.18, issue.3, pp.647-652, 1990.
DOI : 10.1093/nar/18.3.647

URL : https://academic.oup.com/nar/article-pdf/18/3/647/3885916/18-3-647.pdf

H. M. Mahbubani, T. Paull, J. K. Elder, and J. J. Blow, DNA replication initiates at multiple sites on plasmid DNA in Xenopus egg extracts, Nucleic Acids Research, vol.20, issue.7, pp.1457-1462, 1992.
DOI : 10.1093/nar/20.7.1457

Y. Marahrens and B. Stillman, A yeast chromosomal origin of DNA replication defined by multiple functional elements, Science, vol.255, issue.5046, pp.817-823, 1992.
DOI : 10.1126/science.1536007

Y. Marahrens and B. Stillman, Replicator dominance in a eukaryotic chromosome, EMBO J, vol.13, pp.3395-3400, 1994.

L. Martin-parras, P. Hernandez, M. L. Martinez-robles, and J. B. Schvartzman, Unidirectional replication as visualized by two-dimensional agarose gel electrophoresis, Journal of Molecular Biology, vol.220, issue.4, pp.843-852, 1991.
DOI : 10.1016/0022-2836(91)90357-C

M. Matsuoka, M. Matsubara, H. Daidoh, T. Imanaka, K. Uchida et al., Analysis of regions essential for the function of chromosomal replicator sequences from Yarrowia lipolytica, Mol. Gen. Genet, vol.237, pp.327-333, 1993.

K. Maundrell, A. Hutchinson, and S. Shall, Sequence analysis of ARS elements in fission yeast, EMBO J, vol.7, pp.2203-2209, 1988.

T. Nakase and K. Komagata, SIGNIFICANCE OF DNA BASE COMPOSITION IN THE CLASSIFICATION OF YEAST GENUS CANDIDA, The Journal of General and Applied Microbiology, vol.17, issue.3, pp.259-279, 1971.
DOI : 10.2323/jgam.17.259

D. A. Natale, R. M. Umek, and D. Kowalski, Ease of DNA unwinding is a conserved property of yeast replication origins, Nucleic Acids Research, vol.21, issue.3, pp.555-560, 1993.
DOI : 10.1093/nar/21.3.555

C. S. Newlon, I. Collins, A. Dershowitz, A. M. Deshpande, S. A. Greenfeder et al., Analysis of Replication Origin Function on Chromosome III of Saccharomyces cerevisiae, Cold Spring Harbor Symposia on Quantitative Biology, vol.58, issue.0, pp.415-423, 1993.
DOI : 10.1101/SQB.1993.058.01.048

C. S. Newlon and J. F. Theis, The structure and function of yeast ARS elements, Current Opinion in Genetics & Development, vol.3, issue.5, pp.752-758, 1993.
DOI : 10.1016/S0959-437X(05)80094-2

M. Ohkuma, K. Kobayashi, S. Kawai, C. W. Hwang, A. Ohta et al., Identification of a centromeric activity in the autonomously replicating TRA region allows improvement of the host-vector system for Candida maltosa, MGG Molecular & General Genetics, vol.249, issue.4, pp.447-455, 1995.
DOI : 10.1007/BF00287107

T. L. Orr-weaver and J. W. Szostak, Yeast recombination: the association between double-strand gap repair and crossing-over., Proc. Natl. Acad. Sci. USA, pp.4417-4421, 1983.
DOI : 10.1073/pnas.80.14.4417

URL : http://www.pnas.org/content/80/14/4417.full.pdf

J. Pla, R. M. Perez-diaz, F. Navarro-garcia, M. Sanchez, and C. Nombela, Cloning of the Candida albicans HIS1 gene by direct complementation of a C. albicans histidine auxotroph using an improved double-ARS shuttle vector, Gene, vol.165, issue.1, pp.115-120, 1995.
DOI : 10.1016/0378-1119(95)00492-O

H. Rao, Y. Marahrens, and B. Stillman, Functional conservation of multiple elements in yeast chromosomal replicators., Molecular and Cellular Biology, vol.14, issue.11, pp.7643-7651, 1994.
DOI : 10.1128/MCB.14.11.7643

M. Roberge and S. M. Gasser, DNA loops: structural and functional properties of scaffold-attached regions, Molecular Microbiology, vol.264, issue.4, pp.419-423
DOI : 10.1016/0092-8674(90)90071-L

URL : http://onlinelibrary.wiley.com/doi/10.1111/j.1365-2958.1992.tb01485.x/pdf

R. Roggenkamp, H. Hansen, M. Eckart, Z. Janowicz, and C. P. Hollenberg, Transformation of the methylotrophic yeast Hansenula polymorpha by autonomous replication and integration vectors, MGG Molecular & General Genetics, vol.24, issue.2, pp.302-308, 1986.
DOI : 10.1007/BF00331655

M. A. Romanos, C. A. Scorer, and J. J. Clare, Foreign gene expression in yeast: a review, Yeast, vol.261, issue.6, pp.423-448, 1992.
DOI : 10.1128/MCB.4.10.1985

J. Sambrook, E. F. Fritsch, and T. Maniatis, Molecular cloning: a laboratory manual, 1989.

T. Shinomiya and S. Ina, Mapping an initiation region of DNA replication at a single-copy chromosomal locus in Drosophila melanogaster cells by two-dimensional gel methods and PCR-mediated nascent-strand analysis: multiple replication origins in a broad zone., Molecular and Cellular Biology, vol.14, issue.11, pp.7394-7403, 1994.
DOI : 10.1128/MCB.14.11.7394

K. Shirahige, T. Iwasaki, M. B. Rashid, N. Ogasawara, and H. Yoshikawa, Location and characterization of autonomously replicating sequences from chromosome VI of Saccharomyces cerevisiae., Molecular and Cellular Biology, vol.13, issue.8, pp.5043-5056, 1993.
DOI : 10.1128/MCB.13.8.5043

J. G. Smith, M. S. Caddle, G. H. Bulboaca, J. G. Wohlgemuth, M. Baum et al., Replication of centromere II of Schizosaccharomyces pombe., Molecular and Cellular Biology, vol.15, issue.9, pp.5165-5172, 1995.
DOI : 10.1128/MCB.15.9.5165

P. E. Sudbery, The non-Saccharomyces yeasts, Yeast, vol.142, issue.13, pp.1707-1726, 1994.
DOI : 10.1128/MCB.12.5.1977

K. Takahashi, S. Murakami, Y. Chikashige, H. Funabiki, O. Niwa et al., A low copy number central sequence with strict symmetry and unusual chromatin structure in fission yeast centromere., Molecular Biology of the Cell, vol.3, issue.7, pp.819-835, 1992.
DOI : 10.1091/mbc.3.7.819

J. F. Theis and C. S. Newlon, Domain B of ARS307 contains two functional elements and contributes to chromosomal replication origin function., Molecular and Cellular Biology, vol.14, issue.11, pp.7652-7659, 1994.
DOI : 10.1128/MCB.14.11.7652

J. G. Wohlgemuth, G. H. Bulboaca, M. Moghadam, M. S. Caddle, and M. P. Calos, Physical mapping of origins of replication in the fission yeast Schizosaccharomyces pombe., Molecular Biology of the Cell, vol.5, issue.8, pp.839-849, 1994.
DOI : 10.1091/mbc.5.8.839

J. W. Xuan, P. Fournier, and C. Gaillardin, Cloning of the LYS5 gene encoding saccharopine dehydrogenase from the yeast Y. lipolytica by targeted integration, Curr. Genet, vol.14, pp.19-21, 1988.

H. Zhu, C. S. Newlon, and J. A. Huberman, Localization of a DNA replication origin and termination zone on chromosome III of Saccharomyces cerevisiae., Molecular and Cellular Biology, vol.12, issue.10, pp.4733-4741, 1992.
DOI : 10.1128/MCB.12.10.4733

J. G. Zhu, D. L. Carlson, D. D. Dubey, K. Sharma, and J. A. Huberman, Comparison of the two major ARS elements of the ura4 replication origin region with other ARS elements in the fission yeast, Schizosaccharomyces pombe, Chromosoma, vol.102, issue.6, pp.414-422, 1994.
DOI : 10.1007/BF00362286

M. Zimmermann and P. Fournier, Electrophoretic Karyotyping of Yeasts, Nonconventional yeasts in biotechnology?a handbook, pp.101-116, 1996.
DOI : 10.1007/978-3-642-79856-6_3