M. Gilpin, The genetic effective size of a metapopulation, Biol J Linn Soc, vol.42, pp.165-175, 1991.

S. Wright, Evolution and the Genetics of Populations. Variability within and among Natural Populations, vol.4, 1978.

B. K. Epperson, Geographical Genetics Princeton, 2003.

M. Baguette, S. Petit, and F. Queva, Population spatial structure and migration of three butterfly species within the same habitat network: consequences for conservation, J Appl Ecol, vol.37, pp.100-108, 2000.

M. Baguette and H. Van-dyck, Landscape connectivity and animal behavior: functional grain as a key determinant for dispersal, Landscape Ecol, vol.22, pp.117-1129, 2007.

H. B. Britten, P. F. Brussard, D. D. Murphy, and P. R. Ehrlich, A test for isolation-by-distance in central rocky mountain and great basin populations of Edith's checkerspot butterfly (Euphydryas editha), J Hered, vol.86, pp.204-210, 1995.

J. Blondel, D. W. Thomas, A. Charmantier, P. Perret, and P. Bourgault, Lambrechts MM: A thirty-year study of phenotypic and genetic variation of blue tits in Mediterranean habitat mosaics, vol.56, pp.661-673, 2006.

M. Heino and I. Hanski, Evolution of migration rate in a spatially realistic metapopulation model, Am Nat, vol.157, pp.495-511, 2001.

H. Kokko and A. Lopez-sepulcre, From individual dispersal to species ranges: Perspectives for a changing world, Science, vol.313, pp.789-791, 2006.

N. Schtickzelle, G. Mennechez, and M. Baguette, Dispersal depression with habitat fragmentation in the bog fritillary butterfly, Ecology, vol.87, pp.1057-1065

S. Harrison, P. J. Edwards, R. M. May, and N. R. Webb, Metapopulation and Conservation, Large Scale Ecology and Conservation Biology, pp.111-128, 1994.

I. Hanski and . Oxford, , 1999.

M. Slatkin, Isolation by distance in equilibrium and non-equilibrium populations, Evolution, vol.47, pp.264-279, 1993.

R. Nathan, G. Perry, J. T. Cronin, A. E. Strand, and M. L. Cain, Methods for estimating long-distance dispersal, Oikos, vol.103, pp.261-273, 2003.

M. C. Whitlock and D. E. Mccauley, Indirect measures of gene flow and migration: F ST ?1/(4Nm+1), Heredity, vol.82, pp.117-125, 1999.

J. Tufto, S. Engen, and K. Hindar, Inferring patterns of migration from gene frequencies under equilibrium conditions, Genetics, vol.144, pp.1911-1921, 1996.

J. E. Neigel, F. Is, and . St-obsolete, Conserv Genet, vol.3, pp.167-173, 2002.

, Descimon H: L'acclimatation de lépidoptères: un essai d'expérimentation en biogéographie. Alexanor, vol.9, pp.195-204, 1976.

B. Barascud, J. F. Martin, M. Baguette, and H. Descimon, Genetic consequences of an introduction-colonization process in an endangered butterfly species, J Evol Biol, vol.12, pp.697-709, 1999.

H. Descimon, M. Zimmermann, E. Cosson, B. Barascud, and G. Nève, Diversité génétique, variation géographique et flux géniques chez quelques Lépidoptères Rhopalocères français, Gen Sel Evol, vol.33, issue.1, pp.223-249, 2001.

G. Nève, B. Barascud, H. Descimon, and M. Baguette, Genetic structure of Proclossiana eunomia populations at the regional scale (Lepidoptera, Nymphalidae). Heredity, vol.84, pp.657-666, 2000.

A. H. Porter and H. Geiger, Limitations to the inference of gene flow at regional geographic scales -an example from the Pieris napi group (Lepidoptera: Pieridae) in Europe, Biol J Linn Soc, vol.54, pp.329-348, 1995.

G. Nève, Population genetics of butterflies, The Ecology of Butterflies in Europe Edited by: Settele J, Shreeve TG

G. Nève, B. Barascud, R. Hughes, A. J. Descimon, H. Lebrun et al., Dispersal, colonization power and metapopulation structure in the vulnerable butterfly Proclossiana eunomia (Lepidoptera, Nymphalidae), J Appl Ecol, vol.33, pp.14-22, 1996.

D. Louy, J. C. Habel, T. Schmitt, T. Assmann, M. Meyer et al., Strongly diverging population genetic patterns of three skipper species: the role of habitat fragmentation and dispersal ability, Conserv Genet, vol.8, pp.671-681, 2007.

N. Keyghobadi, J. Roland, and C. Strobeck, Genetic differentiation and gene flow among populations of the alpine butterfly, Parnassius smintheus, vary with landscape connectivity, Mol Ecol, vol.14, pp.1897-1909, 2005.

N. Schtickzelle and M. Baguette, Behavioural responses to habitat patch boundaries restrict dispersal and generate emigration-patch area relationship in fragmented landscapes with low quality matrix, J Anim Ecol, vol.72, pp.533-545, 2003.

N. Schtickzelle, A. Joiris, H. Van-dyck, and M. Baguette, Quantitative analysis of changes in movement behaviour within and outside habitat in a specialist butterfly, BMC Evol Biol, vol.7, p.4, 2007.

H. Van-dyck and M. Baguette, The behavioural nature of dispersal in fragmented landscapes: routine or special movements?, Basic App Ecol, vol.6, pp.535-545, 2005.

N. Keyghobadi, K. P. Unger, J. D. Weintraub, and D. M. Fonseca, Remnant populations of the Regal Fritillary (Speyeria idalia) in Pennsylvania: Local genetic structure in a high gene flow species, Conserv Genet, vol.7, pp.309-313, 2006.

C. R. Haag, M. Saastamoinen, J. H. Marden, and I. Hanski, A candidate locus for variation in dispersal rate in a butterfly metapopulation, P Roy Soc B-Biol Sci, vol.272, pp.2449-2456, 2005.

, Bohonak AJ: Dispersal, gene flow and population structure, Q Rev Biol, vol.74, pp.21-45, 1999.

J. Clobert and J. D. Lebreton, Estimation of demographic parameters in bird populations, Bird Populations Studies, Relevance to Conservation and Management Edited by: Perrins CM, Lebreton JD, Hirons GJM, pp.75-104, 1991.

J. Clobert, R. Ims, and F. Rousset, Causes, mechanisms and consequences of dispersal. In Ecology, genetics and evolution of metapopulation Edited by: Hanski I, Gaggiotti OE, pp.307-335, 2004.

M. Baguette, I. Convié, and G. Nève, Male density effect on female spatial behaviour in the butterfly Proclossiana eunomia, Acta OEcol, vol.17, pp.225-232, 1996.

M. Baguette, M. Mennechez, S. Petit, and N. Schtickzelle, Effect of habitat fragmentation on dispersal in the butterfly Proclossiana eunomia, C R Biologies, vol.326, pp.200-209, 2003.

C. Smithers, Handbook of Insect Collecting, Collection, Preparation, Preservation and Storage Newton Abbott: David and Charles, 1982.

V. Nabokov, Nabokov's Butterflies, p.640, 1966.

E. Meglécz, F. Péténian, E. Danchin, A. C. Acier, J. Y. Rasplus et al., High similarity between flanking regions of different microsatellites detected within each of two species of Lepidoptera: Parnassius apollo and Euphydryas aurinia, Mol Ecol, vol.13, pp.1693-1700, 2004.

E. Meglécz, A. Anderson, D. Bourguet, R. Butcher, A. Caldas et al., Microsatellite flanking region similarities among different loci within insect species, Insect Mol Biol, vol.16, pp.175-185, 2007.

B. J. Richardson, P. R. Baverstock, and M. Adams, Allozyme Electrophoresis Sydney, 1986.

F. Rousset and . Genepop, 007: a complete re-implementation of the GENEPOP software for Windows and Linux, Mol Ecol Res, vol.8, pp.103-106, 2008.
URL : https://hal.archives-ouvertes.fr/halsde-00366752

. Sas-institute-inc, User's Guide. Version 6 4th edition, SAS/STAT(R), 1990.

, R Development Core Team: R: A language and environment for statistical computing. Vienna: R Foundation for Statistical Computing, 2007.

S. Piry, A. Alapetite, J. M. Cornuet, D. Paetkau, L. Baudouin et al., GeneClass2: a software for genetic assignment and first-generation migrant detection, J Hered, vol.95, pp.536-539, 2004.

B. Rannala and J. L. Mountain, Detecting immigration by using multilocus genotypes, Proc Natl Acad Sci, vol.94, pp.9197-9221, 1997.

D. Paetkau, R. Slade, M. Burden, and A. Estoup, Genetic assignment methods for the direct, real-time estimation of migration