J. C. Avise, Phylogeography-the history and formation of species, 2000.

G. M. Hewitt, The structure of biodiversity-insights from molecular phylogeography, Front Zool, vol.1, p.15679920, 2004.

G. M. Hewitt, Post-glacial re-colonization of European biota, Biol J Linn Soc Lond, vol.68, pp.87-112, 1999.

J. M. Seddon, F. Santucci, N. J. Reeve, and G. M. Hewitt, DNA footprints of European hedgehogs, Erinaceus europaeus and E. concolor: Pleistocene refugia, postglacial expansion and colonization routes, Mol Ecol, vol.10, p.11555261, 2001.

Z. Varga and T. Schmitt, Types of areal and oreotundral disjunctions in the western Palearctic, Biol J Linn Soc Lond, vol.93, pp.415-430, 2008.

M. Ronikier, G. M. Schneeweiss, and P. Schoönswetter, The extreme disjunction between Beringia and Europe in Ranunculus glacialis s. l. (Ranunculaceae) does not coincide with the deepest genetic split -a story of the importance of temperate mountain ranges in arctic-alpine phylogeography, Mol Ecol, vol.21, p.23057869, 2012.

T. Schmitt and Z. Varga, Extra-Mediterranean refugia: The rule and not the exception?, Front Zool, vol.9, p.22953783, 2012.

G. De-lattin, Grundriss der Zoogeographie, 1967.

T. Schmitt, Molecular biogeography of Europe: Pleistocene cycles and postglacial trends, Front Zool, vol.4, p.17439649, 2007.

L. Després, S. Loriot, and M. Gaudeul, Geographic pattern of genetic variation in the European globeflower Trollius europaeus L. (Ranunculaceae) inferred from amplified fragment length polymorphism markers, Mol Ecol, vol.11, p.12406244, 2002.

K. Kramp, S. Huck, M. Niketi?, G. Tomovi?, and T. Schmitt, Multiple glacial refugia and complex postglacial range shifts of the obligatory woodland plant Polygonatum verticillatum (Convallariaceae), Plant Biol (Stuttg), vol.11, pp.392-404, 2009.

C. Drees, M. Husemann, K. Homburg, P. Brandt, P. Dieker et al., Molecular analyses and species distribution models indicate cryptic northern mountain refugia for a forest-dwelling ground beetle, J Biogeogr, vol.43, pp.2223-2236, 2016.

J. Habel, T. Schmitt, M. Meyer, A. Finger, D. Rödder et al., Biogeography meets conservation: the genetic structure of the endangered lycaenid butterfly Lycaena helle (Denis & Schiffermü ller, 1775), Biol J Linn Soc Lond, vol.101, pp.155-168, 2010.

K. Kramp, O. Cizek, P. M. Madeira, A. A. Ramos, M. Konvicka et al., Genetic implications of phylogeographical patterns in the conservation of the boreal wetland butterfly Colias palaeno

, Biol J Linn Soc Lond, vol.119, pp.1068-1081, 2016.

G. Nève, B. Barascud, R. Hughes, A. J. Descimon, H. Lebrun et al., Dispersal, colonization power and metapopulation structure in the vulnerable butterfly Proclossiana eunomia (Lepidoptera, Nymphalidae), J Appl Ecol, vol.33, pp.14-22, 1996.

H. A. Binney, K. J. Willis, M. E. Edwards, S. A. Bhagwat, P. M. Anderson et al., The distribution of late-Quaternary woody taxa in northern Eurasia: evidence from a new macrofossil database, Quat Sci Rev, vol.28, pp.2445-2464, 2009.

J. Ehlers and P. L. Gibbard, The extent and chronology of Cenozoic global glaciation, Quat Int, vol.164, pp.6-20, 2007.

P. E. Tarasov, V. S. Volkova, T. Webb, . Iii, J. Guiot et al., Last glacial maximum biomes reconstructed from pollen and plant macrofossil data from northern Eurasia, J Biogeogr, vol.27, pp.609-620, 2000.
URL : https://hal.archives-ouvertes.fr/hal-01457720

R. J. Abbott and C. Brochmann, History and evolution of the Arctic flora: in the footsteps of Eric Hultén, Mol Ecol, vol.12, p.12535083, 2003.

V. G. Goetcheus and H. H. Birks, Full-glacial upland tundra vegetation preserved under tephra in the Beringian National Park, Quat Sci Rev, vol.20, pp.135-147, 2001.

I. G. Alsos, T. Engelskjøn, L. Gielly, P. Taberlet, and C. Brochmann, Impact of ice ages on circumpolar molecular diversity: insights from an ecological key species, Mol Ecol, vol.14, p.16029475, 2005.
URL : https://hal.archives-ouvertes.fr/halsde-00273936

P. B. Eidesen, I. G. Alsos, M. Popp, O. Stensrud, J. Suda et al., Nuclear vs. plastid data: complex Pleistocene history of a circumpolar key species, Mol Ecol, vol.16, p.17850553, 2007.

P. Mardulyn, Y. Mikhailov, and J. M. Pasteels, Testing phylogeographic hypotheses in a Euro-Siberian coldadapted leaf beetle with coalescent simulations, Evolution, vol.63, p.19545269, 2009.

G. Nève, B. Barascud, H. Descimon, and M. Baguette, Gene flow rise with habitat fragmentation in the bog fritillary butterfly (Lepidoptera: Nymphalidae), BMC Evol Biol, vol.8, p.18366652, 2008.

G. Nève, A. Pavlí?ko, and M. Konvi?ka, Loss of genetic diversity through spontaneous colonization in the bog fritillary butterfly, Proclossiana eunomia (Lepidoptera: Nymphalidae) in the Czech Republic, Eur J Entomol, vol.106, pp.11-19, 2009.

V. K. Tuzov and G. C. Bozano, Guide to the butterflies of the Palearctic region, Tribe Argynnini: Boloria, Proclossiana, Clossiana. Milano: Omnes Artes, vol.II, 2006.

B. Petersen, Some trends of speciation in the cold-adapted Holarctic fauna, Zoologiska Bidrag från Uppsala, vol.30, pp.233-314, 1956.

P. Jak?i? and C. Van-swaay, Boloria eunomia (Esper, 1799): a new species for Serbia (Nymphalidae), Nota Lepidopterol, vol.30, pp.64-70, 2007.

O. Kudrna, J. Pennerstorfer, and K. Lux, Distribution atlas of European butterflies and skippers. 3 rd ed. Schwanfeld: Wissenschaftlicher Verlag Peks, 2015.

N. Schtickzelle, C. Turlure, and M. Baguette, Grazing management impacts on the viability of the threatened bog fritillary butterfly Proclossiana eunomia, Biol Conserv, vol.136, pp.651-660, 2007.

P. Klimczuk and M. Sielezniew, Unexpected differences in butterfly diversity between two peat bogs in the same area, Pol J Entomol, vol.86, pp.251-273, 2017.

T. Tolman and R. Lewington, Collins butterfly guide: the most complete field guide to the butterflies of Britain and Europe. 3 rd ed. London: HarperCollins UK, 2008.

C. Yui and A. G. Tatarinov, Butterflies (Lepidoptera, Rhopalocera) in the Arctic fauna, Entomol Rev, vol.86, issue.7, pp.760-786, 2006.

J. Bene?, M. Konvika, J. Dvo?ák, Z. Fric, Z. Havelda et al., Butterflies of the Czech Republic: Distribution and Conservation I, 2002.

R. A. Layberry, P. W. Hall, and J. Lafontaine, The Butterflies of Canada, 1998.

V. V. Tshikolovets, Butterflies of Europe & the Mediterranean area. Kiev: Tshikolovets, 2011.

N. Wahlberg and C. W. Wheat, Genomic outposts serve the phylogenomic pioneers: designing novel nuclear markers for genomic DNA extractions of Lepidoptera, Syst Biol, vol.57, p.18398768, 2008.

A. Monteiro and N. E. Pierce, Phylogeny of Bicyclus (Lepidoptera: Nymphalidae) inferred from COI, COII, and EF-1alpha gene sequences, Mol Phylogenet Evol, vol.18, p.11161761, 2001.

M. Kearse, R. Moir, A. Wilson, S. Stones-havas, M. Cheung et al., Geneious Basic: an integrated and extendable desktop software platform for the organization and analysis of sequence data, Bioinformatics, vol.28, p.22543367, 2012.

M. Clement, D. Posada, and K. Crandall, TCS: a computer program to estimate gene genealogies, Mol Ecol, vol.9, p.11050560, 2000.
DOI : 10.1046/j.1365-294x.2000.01020.x

J. W. Leigh and D. Bryant, PopART: full-feature software for haplotype network construction, Methods Ecol Evol, vol.6, pp.1110-1116, 2015.
DOI : 10.1111/2041-210x.12410
URL : https://besjournals.onlinelibrary.wiley.com/doi/pdf/10.1111/2041-210X.12410

S. Dellicour, P. Mardulyn, and . Spads, 0: a toolbox to perform spatial analyses on DNA sequence data sets, Mol Ecol Resour, vol.14, p.24215429, 2014.
DOI : 10.1111/1755-0998.12200

L. Excoffier, G. Laval, and S. Schneider, Arlequin vers. 3.0: an integrated software package for population genetics data analysis, Evol Bioinform Online, vol.1, pp.47-50, 2005.

L. Excoffier and H. Lischer, Arlequin suite ver 3.5: A new series of programs to perform population genetics analyses under Linux and Windows, Mol Ecol Resour, vol.10, p.21565059, 2010.

R. Lanfear, B. Calcott, H. Sy, and S. Guindon, Partition-finder: combined selection of partitioning schemes and substitution models for phylogenetic analyses, Mol Biol Evol, vol.29, p.22319168, 2012.

A. J. Drummond, M. A. Suchard, D. Xie, and A. Rambaut, Bayesian phylogenetics with BEAUti and the BEAST 1.7, Mol Biol Evol, vol.29, p.22367748, 2012.
DOI : 10.1093/molbev/mss075
URL : https://academic.oup.com/mbe/article-pdf/29/8/1969/13648091/mss075.pdf

N. Wahlberg, That awkward age for butterflies: insights from the age of the butterfly subfamily Nymphalinae, Syst Biol, vol.55, p.16952908, 2006.

N. Wahlberg, C. W. Wheat, and C. Peña, Timing and patterns in the taxonomic diversification of Lepidoptera (butterflies and moths), PLoS One, vol.8, issue.11, p.80875, 2013.

T. J. Simonsen, N. Wahlberg, A. D. Warren, and F. Sperling, The evolutionary history of Boloria (Lepidoptera: Nymphalidae): phylogeny, zoogeography and larval foodplant relationships, Syst Biodivers, vol.8, pp.513-529, 2010.

A. J. Drummond and A. Rambaut, BEAST: Bayesian evolutionary analysis by sampling trees, BMC Evol Biol, vol.7, p.17996036, 2007.
DOI : 10.1186/1471-2148-7-214
URL : https://bmcevolbiol.biomedcentral.com/track/pdf/10.1186/1471-2148-7-214

M. A. Miller, W. Pfeiffer, and T. Schwartz, Creating the CIPRES Science Gateway for inference of large phylogenetic trees, Proceedings of the Gateway Computing Environments Workshop (GCE), 2010.

V. V. Tshikolovets, V. O. Bidzilya, and M. I. Golovushkin, The butterflies of Transbaikal Siberia. Kiev, Pardubice: Tshikolovets, 2002.

V. V. Tshikolovets and R. V. Yakovlev, Bá lint Z. The butterflies of Mongolia, 2009.

V. V. Tshikolovets, R. V. Yakovlev, and O. E. Kosterin, The butterflies of Altai

P. Kiev, , 2009.

V. V. Tshikolovets and Y. P. Nekrutenko, The butterflies of Caucasus and Transcaucasia (Armenia, Azerbaijan, Georgia and Russia), 2012.

V. V. Tshikolovets, O. Kosterin, P. Gorbunov, and R. Yakovlev, The Butterflies of Kazakhstan. Kiev, Pardubice: Tshikolovets, 2016.

R. J. Hijmans, S. E. Cameron, J. L. Parra, P. G. Jones, and A. Jarvis, Very high resolution interpolated climate surfaces for global land areas, Int J Climatol, vol.25, pp.1965-1978, 2005.
DOI : 10.1002/joc.1276

S. J. Phillips, M. Dudík, and R. E. Schapire, Maxent software for modeling species niches and distributions (Version 3.4.1), 2018.

R. J. Hijmans, S. Phillips, J. Leathwick, J. Elith, and . Dismo, Species Distribution Modeling

R. J. Hijmans and . Raster, Geographic Data Analysis and Modeling. R package version 2.8-4, 2018.

Y. Fourcade, J. O. Engler, D. Rodder, and J. Secondi, Mapping species distributions with MAXENT using a geographically biased sample of presence data: A performance assessment of methods for correcting sampling bias, PLoS One, vol.9, p.24818607, 2014.

. Qgis-development-team, Geographic information system. Open source geospatial foundation project, 2018.

R. Muscarella, P. J. Galante, M. Soley-guardia, R. A. Boria, J. M. Kass et al., ENMeval: an R package for conducting spatially independent evaluations and estimating optimal model complexity for maxent eco-logical niche models, Methods Ecol Evol, vol.5, pp.1198-1205, 2014.
DOI : 10.1111/2041-210x.12261
URL : https://besjournals.onlinelibrary.wiley.com/doi/pdf/10.1111/2041-210X.12261

Y. Fu, Statistical tests of neutrality of mutations against population growth, hitch-hiking, and background selection, Genetics, vol.147, p.9335623, 1997.

F. Tajima, Statistical method for testing the neutral mutation hypothesis by DNA polymorphism, Genetics, vol.123, p.2513255, 1989.

J. Ehlers, P. L. Gibbard, and P. D. Hughes, Quaternary glaciations-extent and chronology, 2011.
DOI : 10.1016/b978-0-08-100524-8.00003-8

V. Todisco, P. Gratton, D. Cesaroni, and V. Sbordoni, Phylogeography of Parnassius apollo: Hhints on taxonomy and conservation of a vulnerable glacial butterfly invader, Biol J Linn Soc Lond, vol.101, pp.169-183, 2010.

I. Kleckova, M. Cesanek, Z. Fric, and L. Pellissier, Diversification of the cold-adapted butterfly genus Oeneis related to Holarctic biogeography and climatic niche shifts, Mol Phylogenet Evol, vol.92, p.26166775, 2015.

C. Peña, H. Witthauer, I. Kle?ková, Z. Fric, and N. Wahlberg, Adaptive radiations in butterflies: evolutionary history of the genus Erebia (Nymphalidae: Satyrinae), Biol J Linn Soc Lond, vol.116, pp.449-467, 2015.

U. Kodandaramaiah and N. Wahlberg, Phylogeny and biogeography of Coenonympha butterflies (Nymphalidae: Satyrinae)-patterns of colonization in the Holarctic, Syst Entomol, vol.34, pp.315-323, 2009.

U. Kodandaramaiah, M. F. Braby, R. Grund, C. J. Mü-ller, and N. Wahlberg, Phylogenetic relationships, biogeography and diversification of Coenonymphina butterflies (Nymphalidae: Satyrinae): intercontinental dispersal of a southern Gondwanan group, Syst Entomol, vol.43, pp.798-809, 2018.

Z. Varga, Das Prinzip der areal-analytischen Methode in der Zoogeographie und die FaunenelementeEinteilung der europaäischen Tagschmetterlinge (Lepidoptera: Diurna), Acta Biologica Debrecina, vol.14, pp.223-285, 1977.

I. Sanmartin, H. Enghoff, and F. Ronquist, Patterns of animal dispersal, vicariance and diversification in the Holarctic, Biol J Linn Soc Lond, vol.73, pp.345-390, 2001.

S. P. Mullen, Wing pattern evolution and the origins of mimicry among North American admiral butterflies (Nymphalidae: Limenitis), Mol Phylogenet Evol, vol.39, p.16500119, 2006.

U. Kodandaramaiah and N. Wahlberg, Out-of-Africa origin and dispersal-mediated diversification of the butterfly genus Junonia (Nymphalidae: Nymphalinae), J Evol Biol, vol.20, p.17887973, 2007.

C. Peña, S. Nylin, A. Freitas, and N. Wahlberg, Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae), Zool Scr, vol.39, pp.243-258, 2010.

C. Peña, S. Nylin, and N. Wahlberg, The radiation of Satyrini butterflies (Nymphalidae: Satyrinae): a challenge for phylogenetic methods, Zool J Linn Soc, vol.161, pp.64-87, 2011.

J. D. Lafontaine and D. M. Wood, A zoogeographic analysis of the Noctuidae (Lepidoptera) of Beringia, and some inferences about past Beringian habitat, Memoirs of the Entomological Society of Canada, vol.144, pp.109-123, 1988.

K. E. Galbreath and J. A. Cook, Genetic consequences of Pleistocene glaciations for the tundra vole (Microtus oeconomus) in Beringia, Mol Ecol, vol.13, p.14653795, 2004.

A. G. Hope, N. Takebayashi, K. E. Galbreath, S. L. Talbot, J. A. Cook et al., Temporal, spatial and ecological dynamics of speciation among amphi-Beringian small mammals, J Biogeogr, vol.40, pp.415-429, 2013.

B. A. Kohli, V. B. Fedorov, E. Waltari, and J. A. Cook, Phylogeography of a Holarctic rodent (Myodes rutilus): Testing high-latitude biogeographical hypotheses and the dynamics of range shifts, J Biogeogr, vol.42, pp.377-389, 2015.

A. S. Dyke, An outline of North American Deglaciation with emphasis on central and northern Canada, Quaternary Glaciations-Extent and Chronology, Part II, pp.373-424, 2004.

G. E. Beatty and J. Provan, Refugial persistence and postglacial recolonization of North America by the cold-tolerant herbaceous plant Orthilia secunda, Mol Ecol, vol.19, p.20958819, 2010.

J. Godbout, J. P. Jaramillo-correa, J. Beaulieu, and J. Bosquet, A mitochondrial DNA minisatellite reveals the postglacial history of jack pine (Pinus banksiana), a broad-range North American conifer, Mol Ecol, vol.14, p.16156818, 2005.

S. T. Jackson, R. S. Webb, K. H. Anderson, J. T. Overpeck, T. Webb et al., Vegetation and environment in Eastern North America during the Last Glacial Maximum, Quat Sci Rev, vol.19, pp.489-508, 2000.

J. P. Jaramillo-correa, J. Beaulieu, and J. Bousquet, Variation in mitochondrial DNA reveals multiple distant glacial refugia in black spruce (Picea mariana), a transcontinental North American conifer, Mol Ecol, vol.13, p.15315685, 2004.

M. Latutrie, Y. Bergeron, and F. Tremblay, Fine-scale assessment of genetic diversity of trembling aspen in northwestern North America, BMC Evol Biol, vol.16, p.27782797, 2016.

J. A. Lee-yaw, J. T. Irwin, and D. M. Green, Postglacial range expansion from northern refugia by the wood frog, Rana sylvatica, Mol Ecol, vol.17, p.18179428, 2008.

P. Li, M. Li, Y. Shi, Y. Zhao, Y. Wan et al., Phylogeography of North American herbaceous Smilax (Smilacaceae): combined AFLP and cpDNA data support a northern refugium in the Driftless Area

, Am J Bot, vol.100, p.23538874, 2013.

K. C. Rowe, E. J. Heske, P. W. Brown, and K. N. Paige, Surviving the ice: Northern refugia and postglacial recolonization, Proc Natl Acad Sci U S A, vol.101, p.15249673, 2004.

L. A. Lait and P. Hebert, Phylogeographic structure in three North American tent caterpillar species, Lepidoptera: Lasiocampidae): Malacosoma americana, M. californica, and M. disstria, vol.6, p.29576956, 2018.
DOI : 10.7717/peerj.4479
URL : https://doi.org/10.7717/peerj.4479

R. V. Yakovlev, N. A. Shapoval, G. N. Kuftina, A. V. Kulak, and S. V. Kovalev, Notes on the molecular taxonomy of the Proclossiana eunomia complex (Lepidoptera, Nymphalidae: Argynnini): analysis of DNA barcodes, Ukr J Ecol, vol.8, issue.1, pp.222-232, 2018.

M. L. Forister, J. A. Fordyce, and A. M. Shapiro, Geological barriers and restricted gene flow in the holarctic skipper Hesperia comma (Hesperiidae), Mol Ecol, vol.13, p.15488006, 2004.

P. Gratton, M. K. Konopinski, and V. Sbordoni, Pleistocene evolutionary history of the Clouded Apollo (Parnassius mnemosyne): genetic signatures of climate cycles and a 'time-dependent' mitochondrial substitution rate, Mol Ecol, vol.17, p.18986502, 2008.

N. Wahlberg and I. Saccheri, The effects of Pleistocene glaciations on the phylogeography of Melitaea cinxia (Lepidoptera: Nymphalidae), Eur J Entomol, vol.104, pp.675-684, 2007.

A. Bartonova, M. Konvicka, S. Korb, K. Kramp, T. Schmitt et al., Range dynamics of Palaearctic steppe species under glacial cycles: the phylogeography of Proterebia afra (Lepidoptera: Nymphalidae: Satyrinae), Biol J Linn Soc Lond, vol.125, issue.4, pp.867-884, 2018.

J. I. Svendsen, H. Alexanderson, V. I. Astakhov, I. Demidov, J. A. Dowdeswell et al., Late Quaternary ice sheet history of Eurasia, Quat Sci Rev, vol.23, pp.1229-1271, 2004.

K. Mä, . Palä-obiologie-der-pflanzen, and . Jena, Verlag Gustav Fischer; 1953. Phylogeography of two Holarctic butterflies, PLOS ONE, 2019.

C. Brunhoff, N. G. Yoccoz, R. A. Ims, and M. Jaarola, Glacial survival or late glacial colonization? Phylogeography of the root vole (Microtus oeconomus) in north-west Norway, J Biogeogr, vol.33, pp.2136-2144, 2006.

K. Mikkola, J. D. Lafontaine, and V. S. Kononenko, Zoogeography of the Holarctic species of the Noctuidae (Lepidoptera): importance of the Beringian refuge, Entomol Fenn, vol.2, pp.157-173, 1991.

M. C. Quinzin, N. S. Dellicour, S. Svenning, J. Mardulyn, and P. , Glacial survival of trophically linked boreal species in northern Europe, Proc R Soc Lond B Biol Sci, vol.284, 2017.

M. Krzywicki, Fauna Papilionoidea i Hesperioidea (Lepidoptera) Puszczy Bia?owieskiej, Polska Akademia Nauk, vol.25, pp.1-213, 1967.

M. Sielezniew, D. Ponikwicka-tyszko, M. Ratkiewicz, I. Dziekanska, A. Kostro-amnroziak et al., Divergent patterns in the mitochondrial and nuclear diversity of the specialized butterfly Plebejus argus (Lepidoptera: Lycaenidae), Eur J Entomol, vol.108, pp.537-545, 2011.

M. Ratkiewicz and B. Jaroszewicz, Allopatric origins of sympatric forms: the skippers Carterocephalus palaemon palaemon, C. p. tolli and C. silvicolus, Ann Zool Fennici, vol.43, pp.285-94, 2006.

M. Sielezniew, A. Kostro-ambroziak, P. Klimczuk, K. Deoniziak, K. Pa?ka et al., Habitat-related differences in the adult longevity of two ecotypes of a specialized butterfly, J Zool, pp.1-12, 2018.

J. Ehlers, V. Astakhov, P. L. Gibbard, J. Mangerud, and J. I. Svendsen, Middle Pleistocene in Eurasia, Encyclopedia of Quaternary Science. 2 nd ed, pp.172-179, 2013.
DOI : 10.1016/b978-0-444-53643-3.00123-0

T. Schmitt, Biogeographical and evolutionary importance of the European high mountain systems, Front Zool, vol.6, p.19480666, 2009.

G. Nève, B. Barascud, and J. J. Windig, Population biology of Proclossiana eunomia (Nymphalidae): Preliminary results on morphometric and allozyme variation in Belgian and French populations, Nota Lepid, vol.5, pp.3-12, 1994.

V. Dinc?, M. Runquist, M. Nilsson, R. Vila, and . Dispersal, fragmentation, and isolation shape the phylogeography of the European lineages of Polyommatus (Agrodiaetus) ripartii (Lepidoptera: Lycaenidae)

, Biol J Linn Soc Lond, vol.109, pp.817-829, 2013.

J. L. Horreo, M. L. Pelaez, T. Suárez, M. C. Breedveld, H. Benoit et al., evolutionary history and effects of glaciations in a species (Zootoca vivipara) inhabiting multiple biogeographic regions, J Biogeogr, vol.45, pp.1616-1627, 2018.

J. Trettin, S. Agrawal, and J. Heinze, Phylogeography of social polymorphism in a boreo-montane ant, BMC Evol Biol, vol.16, p.27334379, 2016.

M. Vences, J. S. Hauswaldt, S. Steinfartz, O. Rupp, A. Goesmann et al., Radically different phylogeographies and patterns of genetic variation in two European brown frogs, genus Rana, Mol Phylogenet Evol, vol.68, issue.3, p.23632031, 2013.

V. Todisco, P. Gratton, E. V. Zakharov, C. W. Wheat, V. Sbordoni et al., Mitochondrial phylogeography of the Holarctic Parnassius phoebus complex supports a recent refugial model for alpine butterflies, J Biogeogr, vol.39, pp.1058-1072, 2012.

R. H. Hegna, J. A. Galarza, and J. Mappes, Global phylogeography and geographical variation in warning coloration of the wood tiger moth (Parasemia plantaginis), J Biogeogr, vol.42, pp.1469-1481, 2015.

U. Kodandaramaiah, E. Weingartner, N. Janz, L. Dalén, and S. Nylin, Population structure in relation to hostplant ecology and Wolbachia infestation in the comma butterfly, J Evol Biol, vol.24, p.21745252, 2011.

U. Kodandaramaiah, M. Konvicka, T. Tammaru, N. Wahlberg, and K. Gotthard, Phylogeography of the threatened butterfly, the woodland brown Lopinga achine (Nymphalidae: Satyrinae): implications for conservation, J Insect Conserv, vol.16, pp.305-313, 2012.

S. Vandewoestijne, M. Baguette, P. M. Brakefield, and I. J. Saccheri, Phylogeography of Aglais urticae (Lepidoptera) based on DNA sequences of the mitochondrial COI gene and control region, Mol Phylogenet Evol, vol.31, p.15062799, 2004.

M. Konvicka, K. Zimmermann, M. Klimova, V. Hula, and Z. Fric, Inverse link between density and dispersal distance in butterflies: field evidence from six co-occurring species, Popul Ecol, vol.54, pp.91-101, 2012.
DOI : 10.1007/s10144-011-0277-2

G. Mennechez, N. Schtickzelle, and M. Baguette, Metapopulation dynamics of the bog fritillary butterfly: comparison of demographic parameters and dispersal between a continuous and a highly fragmented landscape, Lands Ecol, vol.18, pp.279-291, 2003.

J. C. Habel and T. Schmitt, The burden of genetic diversity, Biol Conserv, vol.147, pp.270-274, 2012.