L. F. Abbott and S. B. Nelson, Synaptic plasticity: Taming the beast, Nature Neuroscience, vol.3, pp.1178-1183, 2000.

J. P. Adelman, J. Maylie, and P. Sah, Small-conductance Ca 2+ -activated K+ channels: Form and function, Annual Review of Physiology, vol.74, pp.245-269, 2012.

C. D. Aizenman and D. J. Linden, Rapid, synaptically driven increases in the intrinsic excitability of cerebellar deep nuclear neurons, Nature Neuroscience, vol.3, pp.109-111, 2000.

S. Armano, P. Rossi, V. Taglietti, and E. Angelo, Long-term potentiation of intrinsic excitability at the mossy fiber-granule cell synapse of rat cerebellum, Journal of Neuroscience, vol.20, pp.5208-5216, 2000.

A. Artola and W. Singer, Long-term potentiation and NMDA receptors in rat visual cortex, Nature, vol.330, pp.649-652, 1987.

B. P. Bean, The action potential in mammalian central neurons, Nature Reviews Neuroscience, vol.8, pp.451-465, 2007.

M. F. Bear, A synaptic basis for memory storage in the cerebral cortex, Proceedings of the National Academy of Sciences of the United States of America, vol.93, pp.13453-13459, 1996.

A. Belmeguenai, E. Hosy, F. Bengtsson, C. M. Pedroarena, C. Piochon et al.,

C. Hansel, Intrinsic plasticity complements long-term potentiation in parallel fiber input gain control in cerebellar Purkinje cells, Journal of Neuroscience, vol.30, pp.13630-13643, 2010.

G. Q. Bi and M. M. Poo, Synaptic modifications in cultured hippocampal neurons: Dependence on spike timing, synaptic strength, and postsynaptic cell type, Journal of Neuroscience, vol.18, pp.10464-10472, 1998.

E. L. Bienenstock, L. N. Cooper, and P. W. Munro, Theory for the development of neuron selectivity: Orientation specificity and binocular interaction in visual cortex, Journal of Neuroscience, vol.2, pp.32-48, 1982.

T. V. Bliss and G. L. Collingridge, A synaptic model of memory: Long-term potentiation in the hippocampus, Nature, vol.361, pp.31-39, 1993.

D. H. Brager and D. Johnston, Plasticity of intrinsic excitability during long-term depression is mediated through mGluR-dependent changes in I(h) in hippocampal CA1 pyramidal neurons, Journal of Neuroscience, vol.27, pp.13926-13937, 2007.

E. Campanac, G. Daoudal, N. Ankri, and D. Debanne, Downregulation of dendritic I(h) in CA1 pyramidal neurons after LTP, Journal of Neuroscience, vol.28, pp.8635-8643, 2008.

E. Campanac and D. Debanne, Spike timing-dependent plasticity: A learning rule for dendritic integration in rat CA1 pyramidal neurons, Journal of Physiology, vol.586, pp.779-793, 2008.
URL : https://hal.archives-ouvertes.fr/hal-01766849

E. Campanac, C. Gasselin, A. Baude, S. Rama, N. Ankri et al., Enhanced intrinsic excitability in basket cells maintains excitatory-inhibitory balance in hippocampal circuits, Neuron, vol.77, pp.712-722, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01774392

X. Chen, L. L. Yuan, C. Zhao, S. G. Birnbaum, A. Frick et al.,

D. Johnston, Deletion of Kv4.2 gene eliminates dendritic A-type K+ current and enhances induction of long-term potentiation in hippocampal CA1 pyramidal neurons, Journal of Neuroscience, vol.26, pp.12143-12151, 2006.

L. N. Cooper and M. F. Bear, The BCM theory of synapse modification at 30: Interaction of theory with experiment, Nature Reviews Neuroscience, vol.13, pp.798-810, 2012.

R. H. Cudmore, L. Fronzaroli-molinieres, P. Giraud, and D. Debanne, Spike-time precision and network synchrony are controlled by the homeostatic regulation of the D-type potassium current, Journal of Neuroscience, vol.30, pp.12885-12895, 2010.
URL : https://hal.archives-ouvertes.fr/hal-01766845

R. H. Cudmore and G. G. Turrigiano, Long-term potentiation of intrinsic excitability in LV visual cortical neurons, Journal of Neurophysiology, vol.92, pp.341-348, 2004.

G. Daoudal, Y. Hanada, and D. Debanne, Bidirectional plasticity of excitatory postsynaptic potential (EPSP)-spike coupling in CA1 hippocampal pyramidal neurons, Proceedings of the National Academy of Sciences of the United States of America, vol.99, pp.14512-14517, 2002.
URL : https://hal.archives-ouvertes.fr/hal-01766854

D. Dasgupta and S. K. Sikdar, Calcium permeable AMPA receptor-dependent long lasting plasticity of intrinsic excitability in fast spiking interneurons of the dentate gyrus decreases inhibition in the granule cell layer, Hippocampus, vol.25, pp.269-285, 2015.

D. Dasgupta and S. K. Sikdar, Heterogeneous network dynamics in an excitatoryinhibitory network model by distinct intrinsic mechanisms in the fast spiking interneurons, Brain Research, vol.1714, pp.27-44, 2019.

D. Debanne, B. H. Gahwiler, and S. M. Thompson, Asynchronous pre-and postsynaptic activity induces associative long-term depression in area CA1 of the rat hippocampus in vitro, Proceedings of the National Academy of Sciences of the United States of America, vol.91, pp.1148-1152, 1994.

D. Debanne, B. H. Gahwiler, and S. M. Thompson, Cooperative interactions in the induction of long-term potentiation and depression of synaptic excitation between hippocampal CA3-CA1 cell pairs in vitro, Proceedings of the National Academy of Sciences of the United States of America, vol.93, pp.11225-11230, 1996.

D. Debanne, B. H. Gahwiler, and S. M. Thompson, Long-term synaptic plasticity between pairs of individual CA3 pyramidal cells in rat hippocampal slice cultures, Journal of Physiology, vol.507, pp.237-247, 1998.

D. Debanne, Y. Inglebert, and M. Russier, Plasticity of intrinsic neuronal excitability, Current Opinion in Neurobiology, vol.54, pp.73-82, 2019.
URL : https://hal.archives-ouvertes.fr/hal-01963474

N. Dehorter, G. Ciceri, G. Bartolini, L. Lim, I. Del-pino et al., Tuning of fast-spiking interneuron properties by an activity-dependent transcriptional switch, Science, vol.349, pp.1216-1220, 2015.

N. S. Desai, T. M. Casimiro, S. M. Gruber, and P. W. Vanderklish, Early postnatal plasticity in neocortex of Fmr1 knockout mice, Journal of Neurophysiology, vol.96, pp.1734-1745, 2006.

N. S. Desai, L. C. Rutherford, and G. G. Turrigiano, Plasticity in the intrinsic excitability of cortical pyramidal neurons, Nature Neuroscience, vol.2, pp.515-520, 1999.

N. S. Desai, L. C. Rutherford, and G. G. Turrigiano, BDNF regulates the intrinsic excitability of cortical neurons, Learning & Memory, vol.6, pp.284-291, 1999.

J. F. Disterhoft, D. A. Coulter, and D. L. Alkon, Conditioning-specific membrane changes of rabbit hippocampal neurons measured in vitro, Proceedings of the National Academy of Sciences of the United States of America, vol.83, pp.2733-2737, 1986.

S. M. Dudek and M. F. Bear, Homosynaptic long-term depression in area CA1 of hippocampus and effects of N-methyl-D-aspartate receptor blockade, Proceedings of the National Academy of Sciences of the United States of America, vol.89, pp.4363-4367, 1992.

M. D. Evans, R. P. Sammons, S. Lebron, A. S. Dumitrescu, T. B. Watkins et al.,

M. S. Grubb, Calcineurin signaling mediates activity-dependent relocation of the axon initial segment, Journal of Neuroscience, vol.33, pp.6950-6963, 2013.

E. S. Faber, A. J. Delaney, and P. Sah, SK channels regulate excitatory synaptic transmission and plasticity in the lateral amygdala, Nature Neuroscience, vol.8, pp.635-641, 2005.

Y. Fan, D. Fricker, D. H. Brager, X. Chen, H. C. Lu et al., Activity-dependent decrease of excitability in rat hippocampal neurons through increases in I(h), Nature Neuroscience, vol.8, pp.1542-1551, 2005.

D. E. Feldman, Timing-based LTP and LTD at vertical inputs to layer II/III pyramidal cells in rat barrel cortex, Neuron, vol.27, pp.45-56, 2000.

D. E. Feldman, The spike-timing dependence of plasticity, Neuron, vol.75, pp.556-571, 2012.

A. Frick, J. Magee, and D. Johnston, LTP is accompanied by an enhanced local excitability of pyramidal neuron dendrites, Nature Neuroscience, vol.7, pp.126-135, 2004.

M. A. Gainey, J. W. Aman, and D. E. Feldman, Rapid disinhibition by adjustment of PV intrinsic excitability during whisker map plasticity in mouse S1, Journal of Neuroscience, vol.38, pp.4749-4761, 2018.

C. Gasselin, Y. Inglebert, N. Ankri, and D. Debanne, Plasticity of intrinsic excitability during LTD is mediated by bidirectional changes in h-channel activity, Scientific Reports, vol.7, p.14418, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01766825

C. Gasselin, Y. Inglebert, and D. Debanne, Homeostatic regulation of h-conductance controls intrinsic excitability and stabilizes the threshold for synaptic modification in CA1 neurons, Journal of Physiology, vol.593, pp.4855-4869, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01766843

P. Gastrein, E. Campanac, C. Gasselin, R. H. Cudmore, A. Bialowas et al.,

D. Debanne, The role of hyperpolarization-activated cationic current in spiketime precision and intrinsic resonance in cortical neurons in vitro, Journal of Physiology, vol.589, pp.3753-3773, 2011.
URL : https://hal.archives-ouvertes.fr/hal-01766841

B. Gustafsson, H. Wigstrom, W. C. Abraham, and Y. Y. Huang, Long-term potentiation in the hippocampus using depolarizing current pulses as the conditioning stimulus to single volley synaptic potentials, Journal of Neuroscience, vol.7, pp.774-780, 1987.

B. E. Herring and R. A. Nicoll, Long-term potentiation: from CaMKII to AMPA receptor trafficking, Annual Review of Physiology, vol.78, pp.351-365, 2016.

D. A. Hoffman and D. Johnston, Downregulation of transient K+ channels in dendrites of hippocampal CA1 pyramidal neurons by activation of PKA and PKC, Journal of Neuroscience, vol.18, pp.3521-3528, 1998.

D. A. Hoffman, J. C. Magee, C. M. Colbert, and D. Johnston, K+ channel regulation of signal propagation in dendrites of hippocampal pyramidal neurons, Nature, vol.387, pp.869-875, 1997.

Y. Y. Huang, E. R. Kandel, L. Varshavsky, E. P. Brandon, M. Qi et al.,

R. Bourtchouladze, A genetic test of the effects of mutations in PKA on mossy fiber LTP and its relation to spatial and contextual learning, Cell, vol.83, pp.1211-1222, 1995.

C. A. Hull, Y. Chu, M. Thanawala, and W. G. Regehr, Hyperpolarization induces a long-term increase in the spontaneous firing rate of cerebellar Golgi cells, Journal of Neuroscience, vol.33, pp.5895-5902, 2013.

J. H. Hyun, K. Eom, K. H. Lee, J. Y. Bae, Y. C. Bae et al.,

S. H. Lee, , 2015.

. Kv1, 2 mediates heterosynaptic modulation of direct cortical synaptic inputs in CA3 pyramidal cells, Journal of Physiology, vol.593, pp.3617-3643

S. L. Jones, M. S. To, and G. J. Stuart, Dendritic small conductance calcium-activated potassium channels activated by action potentials suppress EPSPs and gate spike-timing dependent synaptic plasticity, 2017.

U. R. Karmarkar and D. V. Buonomano, Different forms of homeostatic plasticity are engaged with distinct temporal profiles, European Journal of Neuroscience, vol.23, pp.1575-1584, 2006.

S. R. Kelso, A. H. Ganong, and T. H. Brown, Hebbian synapses in hippocampus, Proceedings of the National Academy of Sciences of the United States of America, vol.83, pp.5326-5330, 1986.

J. Kim, S. C. Jung, A. M. Clemens, R. S. Petralia, and D. A. Hoffman, Regulation of dendritic excitability by activity-dependent trafficking of the A-type K+ channel subunit Kv4.2 in hippocampal neurons, Neuron, vol.54, pp.933-947, 2007.

H. H. Kim, J. M. Park, S. H. Lee, and W. K. Ho, Association of mGluR-dependent LTD of excitatory synapses with endocannabinoid-dependent LTD of inhibitory synapses leads to EPSP to spike potentiation in CA1 pyramidal neurons, Journal of Neuroscience, vol.39, pp.224-237, 2019.

F. Kirchheim, S. Tinnes, C. A. Haas, M. Stegen, and J. Wolfart, Regulation of action potential delays via voltage-gated potassium Kv1.1 channels in dentate granule cells during hippocampal epilepsy, Frontiers in Cellular Neuroscience, vol.7, p.248, 2013.

A. Kirkwood, S. M. Dudek, J. T. Gold, C. D. Aizenman, and M. F. Bear, Common forms of synaptic plasticity in the hippocampus and neocortex in vitro, Science, vol.260, pp.1518-1521, 1993.

H. Kuba, R. Yamada, G. Ishiguro, and R. Adachi, Redistribution of Kv1 and Kv7 enhances neuronal excitability during structural axon initial segment plasticity, Nature Communications, vol.6, p.8815, 2015.

K. P. Lamsa, J. H. Heeroma, P. Somogyi, D. A. Rusakov, and D. M. Kullmann, , 2007.

, Anti-Hebbian long-term potentiation in the hippocampal feedback inhibitory circuit, Science, vol.315, pp.1262-1266

M. E. Larkum, J. J. Zhu, and B. Sakmann, Dendritic mechanisms underlying the coupling of the dendritic with the axonal action potential initiation zone of adult rat layer 5 pyramidal neurons, Journal of Physiology, vol.533, pp.447-466, 2001.

K. X. Li, Y. M. Lu, Z. H. Xu, J. Zhang, J. M. Zhu et al.,

X. M. Li, Neuregulin 1 regulates excitability of fast-spiking neurons through Kv1.1 and acts in epilepsy, Nature Neuroscience, vol.15, pp.267-273, 2011.

J. Lisman, A mechanism for the Hebb and the anti-Hebb processes underlying learning and memory, Proceedings of the National Academy of Sciences of the United States of America, vol.86, pp.9574-9578, 1989.

J. Lisman, R. Yasuda, and S. Raghavachari, Mechanisms of CaMKII action in longterm potentiation, Nature Reviews Neuroscience, vol.13, pp.169-182, 2012.

Y. M. Lu, I. M. Mansuy, E. R. Kandel, and J. Roder, Calcineurin-mediated LTD of GABAergic inhibition underlies the increased excitability of CA1 neurons associated with LTP, Neuron, vol.26, pp.197-205, 2000.

A. Maffei and G. G. Turrigiano, Multiple modes of network homeostasis in visual cortical layer 2/3, Journal of Neuroscience, vol.28, pp.4377-4384, 2008.

J. C. Magee, Dendritic lh normalizes temporal summation in hippocampal CA1 neurons, Nature Neuroscience, vol.2, pp.508-514, 1999.

J. C. Magee, Dendritic integration of excitatory synaptic input, Nature Reviews Neuroscience, vol.1, pp.181-190, 2000.

S. Mahon and S. Charpier, Bidirectional plasticity of intrinsic excitability controls sensory inputs efficiency in layer 5 barrel cortex neurons in vivo, Journal of Neuroscience, vol.32, pp.11377-11389, 2012.

R. C. Malenka, Postsynaptic factors control the duration of synaptic enhancement in area CA1 of the hippocampus, Neuron, vol.6, pp.53-60, 1991.

H. Markram, J. Lubke, M. Frotscher, and B. Sakmann, Regulation of synaptic efficacy by coincidence of postsynaptic APs and EPSPs, Science, vol.275, pp.213-215, 1997.

J. Mellor, R. A. Nicoll, and D. Schmitz, Mediation of hippocampal mossy fiber long-term potentiation by presynaptic Ih channels, Science, vol.295, pp.143-147, 2002.

H. Milshtein-parush, S. Frere, L. Regev, C. Lahav, A. Benbenishty et al.,

I. Slutsky, Sensory deprivation triggers synaptic and intrinsic plasticity in the hippocampus, Cerebral Cortex, vol.27, pp.3457-3470, 2017.

P. Mishra and R. Narayanan, High-conductance states and A-type K+ channels are potential regulators of the conductance-current balance triggered by HCN channels, Journal of Neurophysiology, vol.113, pp.23-43, 2015.

H. Misonou, D. P. Mohapatra, E. W. Park, V. Leung, D. Zhen et al., Regulation of ion channel localization and phosphorylation by neuronal activity, Nature Neuroscience, vol.7, pp.711-718, 2004.

R. M. Mulkey and R. C. Malenka, Mechanisms underlying induction of homosynaptic long-term depression in area CA1 of the hippocampus, Neuron, vol.9, pp.967-975, 1992.

A. B. Nelson, M. Faulstich, S. Moghadam, K. Onori, A. Meredith et al., BK channels are required for multisensory plasticity in the oculomotor system, Neuron, vol.93, pp.211-220, 2017.

A. B. Nelson, C. M. Krispel, C. Sekirnjak, and S. Du-lac, Long-lasting increases in intrinsic excitability triggered by inhibition, Neuron, vol.40, pp.609-620, 2003.

T. J. Ngo-anh, B. L. Bloodgood, M. Lin, B. L. Sabatini, J. Maylie et al., SK channels and NMDA receptors form a Ca 2+ -mediated feedback loop in dendritic spines, Nature Neuroscience, vol.8, pp.642-649, 2005.

M. M. Oh, A. G. Kuo, W. W. Wu, E. A. Sametsky, and J. F. Disterhoft, Watermaze learning enhances excitability of CA1 pyramidal neurons, Journal of Neurophysiology, vol.90, pp.2171-2179, 2003.

G. Ohtsuki and C. Hansel, Synaptic potential and plasticity of an SK2 channel gate regulate spike burst activity in cerebellar purkinje cells. iScience, vol.1, pp.49-54, 2018.

G. Ohtsuki, C. Piochon, J. P. Adelman, and C. Hansel, SK2 channel modulation contributes to compartment-specific dendritic plasticity in cerebellar Purkinje cells, Neuron, vol.75, pp.108-120, 2012.

A. L. Orr, J. E. Hanson, D. Li, A. Klotz, S. Wright et al., Beta-Amyloid inhibits E-S potentiation through suppression of cannabinoid receptor 1-dependent synaptic disinhibition, Neuron, vol.82, pp.1334-1345, 2014.

J. T. Paz, S. Mahon, P. Tiret, S. Genet, B. Delord et al., Multiple forms of activity-dependent intrinsic plasticity in layer V cortical neurones in vivo, Journal of Physiology, vol.587, pp.3189-3205, 2009.

A. C. Penn, C. L. Zhang, F. Georges, L. Royer, C. Breillat et al., Hippocampal LTP and contextual learning require surface diffusion of AMPA receptors, Nature, vol.549, pp.384-388, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01930796

K. F. Raab-graham, P. C. Haddick, Y. N. Jan, and L. Y. Jan, Activity-and mTORdependent suppression of Kv1.1 channel mRNA translation in dendrites, Science, vol.314, pp.144-148, 2006.

S. Rama, M. Zbili, A. Fekete, M. Tapia, M. J. Benitez et al.,

D. Debanne, The role of axonal Kv1 channels in CA3 pyramidal cell excitability, Scientific Reports, vol.7, p.315, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01766826

S. T. Ross and I. Soltesz, Long-term plasticity in interneurons of the dentate gyrus, Proceedings of the National Academy of Sciences of the United States of America, vol.98, pp.8874-8879, 2001.

T. C. Sacktor and J. W. Hell, The genetics of PKMzeta and memory maintenance, Science Signaling, p.10, 2017.

E. Santini, G. J. Quirk, and J. T. Porter, Fear conditioning and extinction differentially modify the intrinsic excitability of infralimbic neurons, Journal of Neuroscience, vol.28, pp.4028-4036, 2008.

B. Santoro, R. A. Piskorowski, P. Pian, L. Hu, H. Liu et al., TRIP8b splice variants form a family of auxiliary subunits that regulate gating and trafficking of HCN channels in the brain, Neuron, vol.62, pp.802-813, 2009.

B. R. Sastry, J. W. Goh, and A. Auyeung, Associative induction of posttetanic and long-term potentiation in CA1 neurons of rat hippocampus, Science, vol.232, pp.988-990, 1986.

B. G. Schreurs, D. Tomsic, P. A. Gusev, and D. L. Alkon, Dendritic excitability microzones and occluded long-term depression after classical conditioning of the rabbit's nictitating membrane response, Journal of Neurophysiology, vol.77, pp.86-92, 1997.

M. M. Shah, Cortical HCN channels: Function, trafficking and plasticity, Journal of Physiology, vol.592, pp.2711-2719, 2014.

M. M. Shah, A. E. Anderson, V. Leung, X. Lin, and D. Johnston, Seizure-induced plasticity of h channels in entorhinal cortical layer III pyramidal neurons, Neuron, vol.44, pp.495-508, 2004.

H. G. Shim, S. S. Jang, D. C. Jang, Y. Jin, W. Chang et al., mGlu1 receptor mediates homeostatic control of intrinsic excitability through Ih in cerebellar Purkinje cells, Journal of Neurophysiology, vol.115, pp.2446-2455, 2016.

H. G. Shim, D. C. Jang, J. Lee, G. Chung, S. Lee et al., Longterm depression of intrinsic excitability accompanied by synaptic depression in cerebellar Purkinje cells, Journal of Neuroscience, vol.37, pp.5659-5669, 2017.

V. Sourdet, M. Russier, G. Daoudal, N. Ankri, and D. Debanne, Long-term enhancement of neuronal excitability and temporal fidelity mediated by metabotropic glutamate receptor subtype 5, Journal of Neuroscience, vol.23, pp.10238-10248, 2003.

N. Spruston, Pyramidal neurons: Dendritic structure and synaptic integration, Nature Reviews Neuroscience, vol.9, pp.206-221, 2008.

Q. Q. Sun, Experience-dependent intrinsic plasticity in interneurons of barrel cortex layer IV, Journal of Neurophysiology, vol.102, pp.2955-2973, 2009.

S. J. Tang, G. Reis, H. Kang, A. C. Gingras, N. Sonenberg et al., A rapamycin-sensitive signaling pathway contributes to long-term synaptic plasticity in the hippocampus, Proceedings of the National Academy of Sciences of the United States of America, vol.99, pp.467-472, 2002.

C. M. Tigaret, S. E. Chamberlain, J. Sadowski, J. Hall, M. C. Ashby et al., Convergent metabotropic signaling pathways inhibit SK channels to promote synaptic plasticity in the hippocampus, Journal of Neuroscience, vol.38, pp.9252-9262, 2018.

H. K. Titley, N. Brunel, and C. Hansel, Toward a neurocentric view of learning, Neuron, vol.95, pp.19-32, 2017.

G. G. Turrigiano and S. B. Nelson, Homeostatic plasticity in the developing nervous system, Nature Reviews Neuroscience, vol.5, pp.97-107, 2004.

Z. Wang, N. L. Xu, C. P. Wu, S. Duan, and M. M. Poo, Bidirectional changes in spatial dendritic integration accompanying long-term synaptic modifications, Neuron, vol.37, pp.463-472, 2003.

D. G. Winder and J. D. Sweatt, Roles of serine/threonine phosphatases in hippocampal synaptic plasticity, Nature Reviews Neuroscience, vol.2, pp.461-474, 2001.

J. Xu, N. Kang, L. Jiang, M. Nedergaard, and J. Kang, Activity-dependent longterm potentiation of intrinsic excitability in hippocampal CA1 pyramidal neurons, Journal of Neuroscience, vol.25, pp.1750-1760, 2005.

R. A. Zalutsky and R. A. Nicoll, Comparison of two forms of long-term potentiation in single hippocampal neurons, Science, vol.248, pp.1619-1624, 1990.

F. Zenke and W. Gerstner, Hebbian plasticity requires compensatory processes on multiple timescales. Philosophical Transactions of the Royal Society of London. Series B, Biological sciences, p.372, 2017.

W. Zhang and D. J. Linden, The other side of the engram: Experience-driven changes in neuronal intrinsic excitability, Nature Reviews Neuroscience, vol.4, pp.885-900, 2003.