J. C. Coulson and B. A. Coulson, Measuring immigration and philopatry in seabirds; recruitment to Black-legged Kittiwake colonies, Ibis, vol.150, pp.288-299, 2008.

T. Coulson, E. A. Catchpole, S. D. Albon, B. J. Morgan, J. M. Pemberton et al., Age, sex, density, winter 995 weather, and population crashes in Soay sheep, Science, vol.292, pp.1528-1531, 2001.

F. Dalerum, B. Shults, and K. E. Kunkel, Estimating sustainable harvest in wolverine populations using logistic regression, Journal of Wildlife Management, vol.72, pp.1125-1132, 2008.

J. E. Diffendorfer, Testing models of source-sink dynamics and balanced dispersal, Oikos, vol.81, pp.417-433, 1000.

B. Doligez and T. Pärt, Estimating fitness consequences of dispersal: a road to 'know-where'? Non-random dispersal and the underestimation of dispersers' fitness, Journal of Animal Ecology, vol.77, pp.1199-1211, 2008.
URL : https://hal.archives-ouvertes.fr/hal-00428100

C. P. Doncaster, J. Clobert, B. Doligez, L. Gustafsson, and E. Danchin, , p.1005, 1997.

, Balanced dispersal between spatially varying local populations: an alternative to the source-sink model, The American Naturalist, vol.150, pp.425-445

M. Efford, Demographic consequences of sex-biased dispersal in a population of brushtail possums, Journal of Animal Ecology, vol.67, pp.503-517, 1998.

M. Frederiksen, J. Lebreton, R. Pradel, R. Choquet, and O. Gimenez, , p.1010, 2014.

, Identifying links between vital rates and environment: a toolbox for the applied ecologist, Journal of Applied Ecology, vol.51, pp.71-81

R. D. Furrer and G. Pasinelli, Empirical evidence for source-sink populations: a review on occurrence, assessments and implications, Biological Reviews, vol.91, pp.782-795, 2016.

M. S. Gaines and L. Mcclenaghan, Dispersal in small mammals, Annual Review of 1015 Ecology and Systematics, vol.11, pp.163-196, 1980.

M. Gamelon, V. Grøtan, A. L. Nilsson, S. Engen, J. W. Hurrell et al., Interactions between demography and environmental effects are important determinants of population dynamics, Science Advances, vol.3, 2017.

B. Gardner, J. Reppucci, M. Lucherini, and J. A. Royle, Spatially explicit inference for open populations: estimating demographic parameters from camera-trap studies, Ecology, vol.91, pp.3376-3383, 2010.

C. Genton, A. Pierre, R. Cristescu, F. Lévréro, S. Gatti et al., How Ebola impacts social dynamics in gorillas: a multistate 1025 modelling approach, Journal of Animal Ecology, vol.84, pp.166-176, 2015.

V. Gervasi, H. Brøseth, E. B. Nilsen, H. Ellegren, Ø. Flagstad et al., Compensatory immigration counteracts contrasting conservation strategies of wolverines (Gulo gulo) within Scandinavia, Biological Conservation, vol.191, pp.632-639, 2015.

B. T. Grenfell, O. F. Price, S. D. Albon, and T. H. Clutton-brock, , p.1030, 1992.

, Overcompensation and Population-Cycles in an Ungulate, Nature, vol.355, pp.823-826

V. Grosbois and G. Tavecchia, Modeling dispersal with capture-recapture data: disentangling decisions of leaving and settlement, Ecology, vol.84, pp.1225-1236, 2003.

V. Grøtan, B. Saether, S. Engen, J. H. Van-balen, A. C. Perdeck et al., Spatial and temporal variation in the relative contribution of density dependence, 1035 climate variation and migration to fluctuations in the size of great tit populations, Journal of Animal Ecology, vol.78, pp.447-459, 2009.

I. Hanski, Metapopulation Ecology, p.44, 1999.

I. Hanski, Spatially realistic theory of metapopulation ecology, Naturwissenschaften, vol.88, pp.372-381, 2001.

B. Hansson, S. Bensch, and D. Hasselquist, Lifetime Fitness of Short-and Long-Distance Dispersing Great Reed Warblers, Evolution, vol.58, pp.2546-2557, 2004.

J. P. Haskell, M. E. Ritchie, and H. Olff, Fractal geometry predicts varying body size scaling relationships for mammal and bird home ranges, Nature, vol.418, pp.527-530, 2002.

V. Henaux, T. Bregnballe, and J. Lebreton, Dispersal and recruitment during 1045 population growth in a colonial bird, the great cormorant Phalacrocorax carbo sinensis, JOURNAL OF AVIAN BIOLOGY, vol.38, pp.44-57, 2007.

S. Herrando-perez, S. Delean, B. W. Brook, and C. J. Bradshaw, Density dependence : an ecological Tower of Babel, Oecologia, vol.170, pp.585-603, 2012.

A. Hixon, S. W. Pacala, and S. A. Sandin, Population regulation: historical context 1050 and contemporary challenges of open versus closed systems, Ecology, vol.83, pp.1490-1508, 2002.

K. A. Hobson, Using stable isotopes to trace long-distance dispersal in birds and other taxa, Diversity and Distributions, vol.11, pp.157-164, 2005.

K. A. Hobson and D. R. Norris, Animal migration: a context for using new techniques and approaches, Tracking animal migration with stable isotopes, pp.1-19, 2008.

M. Hörnell-willebrand, T. Willebrand, and A. A. Smith, Seasonal Movements and Dispersal Patterns: Implications for Recruitment and Management of Willow Ptarmigan (Lagopus lagopus), Journal of Wildlife Management, vol.78, pp.194-201, 2014.

D. Jenkins, A. Watson, and G. R. Miller, Population Studies on Red Grouse, p.1060, 1963.

, Lagopus lagopus scoticus (Lath.) in North-East Scotland, Journal of Animal Ecology, vol.32, pp.317-376

G. M. Jolly, Explicit estimates from capture-recapture data with both death and immigration -Stochastic model, Biometrika, vol.52, pp.225-247, 1965.

R. Julliard, P. Perret, and J. Blondel, Reproductive strategies of philopatric and 1065 immigrant blue tits, Acta Oecologia, vol.17, pp.487-501, 1996.

M. Kahlenberg, M. E. Thompson, M. N. Muller, and R. W. Wrangham, Immigration costs for female chimpanzees and male protection as an immigrant counterstrategy to intrasexual aggression, Animal Behaviour, vol.76, pp.1497-1509, 2008.

R. Kays, M. C. Crofoot, W. Jetz, and M. Wikelski, Terrestrial animal tracking as an 1070 eye on life and planet, Science, vol.348, 2015.

L. F. Keller, K. J. Je, M. A. Beaumont, W. M. Hochachka, J. N. Smith et al., Immigration and the ephemerality of a natural population bottleneck : evidence from molecular markers, Proceedings of the Royal Society B-Biological Sciences, vol.268, pp.1387-1394, 2001.

U. Kormann, F. Gugerli, N. Ray, L. Excoffier, and K. Bollmann, Parsimonybased pedigree analysis and individual-based landscape genetics suggest topography to restrict dispersal and connectivity in the endangered capercaillie, Biological Conservation, vol.152, pp.241-252, 2012.

C. J. Krebs, Ecology: the experimental analysis of distribution of abundance 5th Ed, p.1080, 2001.

B. Cummings,

C. J. Krebs, B. L. Keller, and R. H. Tamarin, Microtus population biology : demographic changes in fluctuating populations of M. ochrogaster and M. Pennsylvanicus in Southern Indiana, vol.50, pp.587-607, 1969.

M. P. Kreuzer and N. J. Huntly, Habitat-specific demography: evidence for source-1085 sink population structure in a mammal, the pika, Oecologia, vol.134, pp.343-349, 2003.

D. Lack, The natural regulation of animal numbers, 1954.

X. Lambin, J. Aars, and S. B. Piertney, Dispersal, intraspecific competition, kin competition and kin facilitation: a review of the empirical evidence, 2001.

E. Clobert, A. A. Danchin, . Dhondt-&-j, and . Nichols, , pp.110-122

J. Lebreton, J. D. Nichols, R. J. Barker, R. Pradel, and J. A. Spendelow, Modeling individual animal histories with multistate capture-recapture models, Advances in Ecological Research, vol.41, pp.87-173, 2009.

R. Levins, Some demographic and genetic consequences of environmental heterogeneity for biological control, Bulletin of the Entomological Society of America, vol.15, pp.237-240, 1969.

N. Lieury, A. Besnard, C. Ponchon, A. Ravayrol, and A. Millon, Geographically isolated but demographically connected: Immigration supports efficient conservation 1100 actions in the recovery of a range-margin population of the Bonelli's eagle in France, Biological Conservation, vol.195, pp.272-278, 2016.

N. Lieury, M. Gallardo, C. Ponchon, A. Besnard, and A. Millon, Relative contribution of local demography and immigration in the recovery of a geographicallyisolated population of the endangered Egyptian vulture, Biological Conservation, vol.191, pp.1105-349, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01444831

N. Lieury, S. Ruette, S. Devillard, M. Albaret, F. Drouyer et al., Compensatory immigration challenges predator control: An experimental evidence-based approach improves management, Journal of Wildlife Management, vol.79, pp.425-434, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01445199

A. J. Loveridge, A. W. Searle, F. Murindagomo, and D. W. Macdonald, The impact of sport-hunting on the population dynamics of an African lion population in a protected area, Biological Conservation, vol.4, pp.548-558, 2007.

R. Macarthur and E. O. Wilson, The Theory of Island Biogeography, 1967.

S. Manel, O. E. Gaggiotti, and R. S. Waples, Assignment methods: Matching biological questions with appropriate techniques, Trends in Ecology and Evolution, vol.20, pp.136-142, 2005.
URL : https://hal.archives-ouvertes.fr/halsde-00276308

E. Matthysen, F. Adriaensen, and A. A. Dhondt, Local recruitment of great and blue tits ( Parus major, P. caeruleus ) in relation to study plot size and degree of 1120 isolation, Ecography, vol.24, pp.33-42, 2001.

M. A. Mcpeek and R. D. Holt, The evolution of dispersal in spatially and temporally varying environments, The American Naturalist, vol.140, pp.1010-1027, 1992.

Y. Melero, M. K. Oliver, and X. Lambin, Relationship type affects the reliability of dispersal distance estimated using pedigree inferences in partially sampled populations: 1125 A case study involving invasive American mink in Scotland, Molecular Ecology, vol.26, pp.4059-4071, 2017.

A. Millon, S. J. Petty, and X. Lambin, Pulsed resources affect the timing of first breeding and lifetime reproductive success of tawny owls, Journal of Animal Ecology, vol.79, pp.426-435, 2010.

A. Millon, S. J. Petty, B. Little, O. Gimenez, T. Cornulier et al., Dampening prey cycle overrides the impact of climate change on predator population dynamics: a long-term demographic study on tawny owls, Global Change Biology, vol.20, pp.1770-1781, 2014.

J. O. Milner, E. B. Nilsen, and H. P. Andreassen, Demographic Side Effects of 1135, 2007.

, Selective Hunting in Ungulates and Carnivores, Conservation Biology, pp.36-47

J. D. Nichols, J. E. Hines, J. D. Lebreton, and R. Pradel, Estimation of contributions to population growth: A reverse-time capture-recapture approach, Ecology, vol.81, pp.3362-3376, 2000.
URL : https://hal.archives-ouvertes.fr/hal-02126409

J. D. Nichols and K. H. Pollock, Estimation of recruitment from immigration versus in 1140 situ reproduction using Pollock's robust design, Ecology, vol.71, pp.21-26, 1990.

A. J. Norman and G. Spong, Single nucleotide polymorphism-based dispersal estimates using noninvasive sampling, Ecology and Evolution, vol.5, pp.3056-3065, 2015.

R. B. O'hara, S. Lampila, and M. Orell, Estimation of rates of births, deaths, and immigration from mark-recapture data, Biometrics, vol.65, pp.275-281, 2009.

M. K. Oliver, S. B. Piertney, A. Zalewski, Y. Melero, and X. Lambin, The compensatory potential of increased immigration following intensive American mink population control is diluted by male-biased dispersal, Biological Invasions, vol.18, pp.3047-3061, 2016.

H. H. Van-oosten, C. Van-turnhout, C. A. Hallmann, F. Majoor, and M. Roodbergen, , p.1150

H. Schekkerman, R. Versluijs, S. Waasdorp, and H. Siepel, Site-specific dynamics in remnant populations of Northern Wheatears Oenanthe oenanthe in the Netherlands, Ibis, vol.157, pp.91-102, 2015.

V. M. Pakanen, A. Rönkä, E. J. Belda, A. Luukkonen, L. Kvist et al., Impact of dispersal status on estimates of local population growth rates in a Temminck's 1155 stint Calidris temminckii population, Oikos, vol.119, pp.1493-1503, 2010.

P. J. Palsbøll, Genetic tagging: contemporary molecular ecology, Biological Journal of the Linnean Society, vol.68, pp.3-22, 1999.

M. M. Peacock and A. T. Smith, The Effect of Habitat Fragmentation on Dispersal Patterns, Mating Behavior, and Genetic Variation in a Pika, p.1160, 1997.

, Metapopulation. Oecologia, vol.112, pp.524-533

M. Z. Peery, B. H. Becker, and S. R. Beissinger, Combining demographic and countbased approaches to identify source-sink dynamics of a threatened seabird, Ecological Applications, vol.16, pp.1516-1528, 2006.

M. Z. Peery, S. R. Beissinger, R. F. House, M. Bérubé, L. A. Hall et al., Characterizing source-sink dynamics with genetic parentage assignments, Ecology, vol.89, pp.2746-2759, 2008.

J. M. Pemberton, Wild pedigrees: the way forward, Proceedings of the Royal Society B: Biological Sciences, vol.275, pp.613-621, 2008.

G. Péron, Compensation and additivity of anthropogenic mortality : life-history 1170 effects and review of methods, Journal of Animal Ecology, vol.82, pp.408-417, 2013.

C. M. Perrins, Population Fluctuations and Clutch-Size in the Great Tit, Parus major L, Journal of Animal Ecology, vol.34, pp.601-647, 1965.

M. Van-de-pol, Y. Vindenes, B. Saether, S. Engen, B. J. Ens et al., Effects of climate change and variability on population 1175 dynamics in a long-lived shorebird, Ecology, vol.91, pp.1192-1204, 2010.

K. H. Pollock, A capture-recapture design robust to unequal probability of capture, Journal of Wildlife Management, vol.46, pp.752-757, 1982.

E. Postma and A. J. Van-noordwijk, Gene flow maintains a large genetic difference in clutch size at a small spatial scale, Nature, vol.433, pp.65-68, 2005.

R. Pradel, Utilization of capture-mark-recapture for the study of recruitment and population growth rate, Biometrics, vol.52, pp.371-377, 1996.
URL : https://hal.archives-ouvertes.fr/hal-02126443

H. R. Pulliam, Sources, sinks, and population regulation, The American Naturalist, vol.132, pp.652-661, 1988.

A. S. Pullin and G. B. Stewart, Guidelines for systematic review in conservation and 1185 environmental management, Conservation Biology, vol.20, pp.1647-1656, 2006.

T. V. Riecke, P. J. Williams, T. L. Behnke, D. Gibson, A. G. Leach et al., Integrated population models: Model assumptions and inference, 2019.

E. P. Robertson, R. J. Fletcher, C. E. Cattau, B. J. Udell, B. E. Reichert et al.,

&. Valle and D. , Isolating the roles of movement and reproduction on effective connectivity alters conservation priorities for an endangered bird, Proceedings of the, 2018.

H. S. Robinson, R. B. Wielgus, H. S. Cooley, and S. W. Cooley, Sink populations in carnivore management: cougar demography and immigration in a hunted Population, 2008.

, Ecological Applications, vol.18, pp.1028-1037

L. A. Rollins, L. E. Browning, C. E. Holleley, J. L. Savage, A. F. Russell et al., Building genetic networks using relatedness information: A novel approach for the estimation of dispersal and characterization of group structure in social animals, Molecular Ecology, vol.21, pp.1727-1740, 2012.

J. A. Royle, R. B. Chandler, R. Sollmann, and B. Gardner, Spatial capturerecapture, 2014.

J. P. Runge, M. C. Runge, and J. D. Nichols, The role of local populations within a landscape context: Defining and classifying sources and sinks, The American Naturalist, vol.167, pp.925-938, 2006.

C. S. Rushing, J. A. Hostetler, T. S. Sillett, P. P. Marra, and J. A. Rotenberg, , p.51, 2017.

, Spatial and temporal drivers of avian population dynamics across the annual cycle, Ecology, vol.98, pp.2837-2850

B. Saether, T. H. Ringsby, O. Bakke, and E. J. Solberg, Spatial and temporal variation in demography of a house sparrow metapopulation, Journal of Animal Ecology, vol.68, pp.628-637, 1210.

B. Saether, V. Grøtan, S. Engen, T. Coulson, P. R. Grant et al., Demographic routes to variability and regulation in bird populations, Nature communications, vol.7, p.12001, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01357674

S. Santoro, A. J. Green, and J. Figuerola, Immigration enhances fast growth of a newly-established source population, Ecology, vol.97, pp.1048-1057, 2016.

M. Schaub, A. Aebischer, O. Gimenez, S. Berger, and R. Arlettaz, Massive immigration balances high anthropogenic mortality in a stable eagle owl population: Lessons for conservation, Biological Conservation, vol.143, pp.1911-1918, 2010.

M. Schaub and D. Fletcher, Estimating immigration using a Bayesian integrated population model: choice of parametrization and priors, Environmental and Ecological Statistics, vol.22, pp.535-549, 2015.

M. Schaub, J. Von-hirschheydt, and M. U. Grüebler, Differential contribution of demographic rate synchrony to population synchrony in barn swallows, Journal, p.1225, 2015.

, Animal Ecology, vol.84, pp.1530-1541

M. Schaub, H. Jakober, and W. Stauber, Strong contribution of immigration to local population regulation: Evidence from a migratory passerine, Ecology, vol.94, pp.1828-1838, 2013.

M. Schaub, T. Reichlin, F. Abadi, M. Kéry, L. Jenni et al., The demographic drivers of local population dynamics in two rare migratory birds, 2012.

, Oecologia, vol.168, pp.97-108

M. Schaub and J. A. Royle, Estimating true instead of apparent survival using spatial Cormack-Jolly-Seber models, Methods in Ecology and Evolution, vol.5, pp.1316-1326, 2014.

M. Schaub, B. Ullrich, G. Knötzsch, P. Albrecht, and C. Meisser, Local population dynamics and the impact of scale and isolation: a study on different little owl 1235 populations, Oikos, vol.115, pp.389-400, 2006.

C. J. Schwarz and A. N. Arnason, A general methodology for the analysis of capturerecapture experiments in open populations, Biometrics, vol.52, pp.860-873, 1996.

G. A. Seber, A note on the multiple recapture census, Biometrika, vol.52, pp.249-259, 1965.

S. P. Sharp, M. Simeoni, and B. J. Hatchwell, Dispersal of sibling coalitions promotes 1240 helping among immigrants in a cooperatively breeding bird, Proceedings of the Royal Society B-Biological Sciences, vol.275, pp.2125-2130, 2008.

J. N. Smith, M. J. Taitt, C. M. Rogers, P. Arcese, L. F. Keller et al., A metapopulation approach to the population biology of the song sparrow Melospiza melodia, Ibis, vol.138, pp.120-128, 1996.

J. A. Spendelow, J. D. Nichols, I. C. Nisbet, H. Hays, G. D. Cormons et al., Estimating annual survival and movement rates of adults within a metapopulation of roseate terns, Ecology, vol.76, pp.2415-2428, 1995.

B. Stacey and M. Taper, Environmental variation and the persistence of small populations, Ecological Applications, vol.2, pp.18-29, 1992.

K. L. Szostek, M. Schaub, and P. H. Becker, Immigrants are attracted by local prebreeders and recruits in a seabird colony, Journal of Animal Ecology, vol.83, pp.1015-1024, 2014.

F. H. Tattersall, D. W. Macdonald, B. J. Hart, and W. Manley, Balanced dispersal or source-sink-do both models describe wood mice in farmed landscapes, p.53, 2004.

, Oikos, vol.106, pp.536-550

S. Telfer, S. B. Piertney, J. F. Dallas, W. A. Stewart, F. Marshall et al.,

X. Lambin, Parentage assignment detects frequent and large-scale dispersal in water voles, Molecular Ecology, vol.12, pp.1939-1949, 2003.

D. J. Tempel, M. Z. Peery, and R. J. Gutiérrez, Using integrated population models to improve conservation monitoring : California spotted owls as a case study, Ecological 1260 Modelling, vol.289, pp.86-95, 2014.

S. Tenan, M. Fasola, S. Volponi, and G. Tavecchia, Conspecific and not performance-based attraction on immigrants drives colony growth in a waterbird, Journal of Animal Ecology, vol.86, pp.1074-1081, 2017.

C. D. Thomas and W. E. Kunin, The spatial structure of populations, Journal of Animal, vol.68, pp.647-657, 1265.

A. Tringali and R. Bowman, Suburban immigrants to wildlands disrupt honest signaling in ultraviolet plumage, Avian Conservation and Ecology, vol.10, p.9, 2015.

J. Wang and A. W. Santure, Parentage and sibship inference from multilocus genotype data under polygamy, Genetics, vol.181, pp.1579-1594, 2009.

P. M. Waser and J. D. Hadfield, How much can parentage analyses tell us about precapture dispersal?, Molecular Ecology, vol.20, pp.1277-1288, 2011.

A. R. Watkinson and W. J. Sutherland, Sources, sinks and pseudo-sinks, Journal of Animal Ecology, vol.64, pp.126-130, 1995.

K. Z. Weinbaum, J. S. Brashares, C. D. Golden, and W. M. Getz, Searching for 1275 sustainability: are assessments of wildlife harvests behind the times?, Ecology Letters, vol.16, pp.99-111, 2013.

Z. Wen, J. D. Nichols, K. H. Pollock, and P. M. Waser, , 2014.