A. R. Kleinhesselink, S. M. Magnoli, and J. H. Cushman, Shrubs as ecosystem engineers across an environmental gradient: effects on species richness and exotic plant invasion, Oecologia, vol.175, pp.1277-1290, 2014.

J. P. Wright and C. G. Jones, The concept of organisms as ecosystem engineers ten years on: progress, limitations, and challenges, Bioscience, vol.56, pp.203-209, 2006.

C. G. Jones, J. H. Lawton, and M. Shachak, Positive and negative effects of organisms as physical ecosystem engineers, Ecology, vol.78, pp.1946-1957, 1997.

O. Reichman and E. W. Seabloom, Ecosystem engineering: a trivialized concept? Response from Reichmann and Seabloom, Trends Ecol. Evol, vol.17, 2002.

S. T. Meyer, Ecosystem engineering in fragmented forests: edge-mediated hyperabundance of leaf-cutting ants and resulting impacts on forest structure, microclimate and regeneration, 2008.

A. Hastings, J. E. Byers, J. A. Crooks, K. Cuddington, C. G. Jones et al., , 2007.

, Ecosystem engineering in space and time, Ecol. Lett, vol.10, pp.153-164

C. J. Lortie, R. W. Brooker, P. Choler, Z. Kikvidze, R. Michalet et al., Rethinking plant community theory, Oikos, vol.107, pp.433-438, 2004.
URL : https://hal.archives-ouvertes.fr/halsde-00276315

B. Herault and D. Thoen, How habitat area, local and regional factors shape plant assemblages in isolated closed depressions, Acta Oecologica, vol.35, pp.385-392, 2009.
URL : https://hal.archives-ouvertes.fr/hal-01032144

S. Gámez-virués, Landscape simplification filters species traits and drives biotic homogenization, Nat. Commun, vol.6, pp.1-8, 2015.

J. Skelton, K. M. Geyer, J. T. Lennon, R. P. Creed, and B. L. Brown, Multi-scale ecological filters shape the crayfish microbiome, Symbiosis, vol.72, pp.159-170, 2017.

J. Bremner, Species' traits and ecological functioning in marine conservation and management, J. Exp. Mar. Bio. Ecol, vol.366, pp.37-47, 2008.

N. Poff, Landscape filters and species traits: towards mechanistic understanding and prediction in stream ecology, J. North Am. Benthol. Soc, vol.16, pp.391-409, 1997.

S. Lengyel, A. D. Gove, A. M. Latimer, J. D. Majer, and R. R. Dunn, Convergent evolution of seed dispersal by ants, and phylogeny and biogeography in flowering plants: a global survey, Perspect. Plant Ecol. Evol. Syst, vol.12, pp.43-55, 2010.

D. R. Lane and H. Bassirirad, Diminishing effects of ant mounds on soil heterogeneity across a chronosequence of prairie restoration sites. Pedobiologia (Jena), vol.49, pp.359-366, 2005.

J. Moya-laraño and D. H. Wise, Direct and indirect effects of ants on a forest-floor food web, Ecology, vol.88, pp.1454-1465, 2007.

A. M. Boulton and K. D. Amberman, How ant nests increase soil biota richness and abundance: a field experiment, Biodivers. Conserv, vol.15, pp.69-82, 2006.

S. Schuch, C. Platner, and D. Sanders, Potential positive effect of the ant species Lasius niger on linyphiid spiders, J. Appl. Entomol, vol.132, pp.375-381, 2008.

D. Sanders, M. Schaefer, C. Platner, and G. Griffiths, Intraguild interactions among generalist predator functional groups drive impact on herbivore and decomposer prey, Oikos, vol.120, pp.418-426, 2011.

N. V. Coggan, M. W. Hayward, and H. Gibb, A global database and 'state of the field' review of research into ecosystem engineering by land animals, J. Anim. Ecol, vol.87, pp.974-994, 2018.

D. Sanders and F. Van-veen, Ecosystem engineering and predation: the multi-trophic impact of two ant species, J. Anim. Ecol, vol.80, pp.569-576, 2011.

K. N. Laland, F. J. Odling-smee, and M. W. Feldman, Evolutionary consequences of niche construction and their implications for ecology, Proc. Natl Acad. Sci. USA 96, vol.10, pp.242-252, 1999.

F. M. Azcárate and B. Peco, Harvester ants (Messor barbarus) as disturbance agents in Mediterranean grasslands, J. Veg. Sci, vol.18, pp.103-110, 2007.

T. De-almeida, O. Blight, F. Mesléard, A. Bulot, E. Provost et al., Harvester ants as ecological engineers for Mediterranean grassland restoration: impacts on soil and vegetation, Biol. Conserv, vol.245, 2020.
URL : https://hal.archives-ouvertes.fr/hal-02611640

F. Mesléard, N. Yavercovski, G. Lefebvre, L. Willm, and A. Bonis, High stocking density controls Phillyrea angustifolia in Mediterranean grasslands, Environ. Manage, vol.59, pp.455-463, 2017.

F. Bensettiti, V. Boulet, C. Chavaudret-laborie, and J. Deniaud, Cahiers d'habitats' Natura, Connaissance et gestion des habitats et des espèces d'intérêt communautaire, vol.2, pp.227-229, 2000.

J. M. Blanco-moreno, P. R. Westerman, V. Atanackovic, and J. Torra, The spatial distribution of nests of the harvester ant Messor barbarus in dryland cereals, Insectes Soc, vol.61, pp.145-152, 2014.

P. Cerdan, Etude de la biologie, de l'écologie et du comportement des fourmis moissonneuses du genre Messor (Hymenoptera, Formicidae) en Crau, 1989.

B. Baraibar, J. Torra, and P. R. Westerman, Harvester ant (Messor barbarus (L.)) density as related to soil properties, topography and management in semiarid cereals, Appl. Soil Ecol, vol.51, pp.60-65, 2011.

D. Baize, Guide des analyses en pédologie: 3e édition revue et augmentée, 2018.

S. R. Olsen, C. V. Cole, F. S. Watanabe, and L. A. Dean, Estimation of available phosphorus in soils by extraction with sodium bicarbonate. Circular 939, 1954.

J. Braun-blanquet, N. Roussine, R. Nègre, and L. Emberger, Les groupements végétaux de la France méditerranéenne, 1952.

J. Mamarot, Mauvaises herbes des cultures, 2002.

G. Agati, Z. G. Cerovic, P. Pinelli, and M. Tattini, Lightinduced accumulation of ortho-dihydroxylated flavonoids as non-destructively monitored by chlorophyll fluorescence excitation techniques, Environ. Exp. Bot, vol.73, pp.3-9, 2011.

Z. G. Cerovic, N. Moise, G. Agati, G. Latouche, N. Ben-ghozlen et al., New portable optical sensors for the assessment of winegrape phenolic maturity based on berry fluorescence, J. Food Compos. Anal, vol.21, pp.650-654, 2008.

A. Macfadyen, Improved funnel-type extractors for soil arthropods, J. Anim. Ecol, vol.30, pp.171-184, 1961.

H. Gisin, Ökologie und lebensgemeinschaften der collembolen im schweizer exkursionsgebiet basels, Rev. Suisse Zool, vol.50, pp.131-224, 1943.

K. R. Barker, . Barker, . Barker, . Carter, and . Sasser, Nematode extraction and bioassays, An advance treatise on meloidogyne, vol.2, 1985.

G. W. Yeates, T. Bongers, R. De-goede, D. W. Freckman, and S. S. Georgieva, Feeding habits in soil nematode families and genera-an outline for soil ecologists, J. Nematol, vol.25, pp.315-331, 1993.

D. Bates, M. Mächler, B. M. Bolker, and S. C. Walker, Fitting linear mixed-effects models using lme4, 2015.

, J. Stat. Softw, vol.67, pp.1-48

B. Ripley, B. Venables, D. M. Bates, K. Hornik, A. Gebhardt et al., Package 'MASS.' Support Functions and Datasets for Venables and Ripley's MASS, 2018.

M. E. Brooks, K. Kristensen, K. J. Van-benthem, A. Magnusson, C. W. Berg et al., 2017 glmmTMB balances speed and flexibility among packages for zeroinflated generalized linear mixed modeling, J, vol.9, pp.378-400

J. Cohen, A power primer, Psychol. Bull, vol.112, pp.155-159, 1992.

M. Torchiano, Package 'effsize ': efficient effect size computation, 2019.

C. O. Fritz, P. E. Morris, and J. J. Richler, Effect size estimates: current use, calculations, and interpretation, J. Exp. Psychol. Gen, vol.141, 2012.

A. J. Oksanen, F. G. Blanchet, R. Kindt, P. Legendre, and R. O'hara, Vegan: community ecology package. R Package 2.3-3, J. Veg. Sci, vol.14, pp.927-930, 2016.

Y. Rosseel, Lavaan: an R package for structural equation modeling, J. Stat. Softw, vol.48, pp.1-36, 2012.

J. B. Grace, Structural equation modeling and natural systems, 2006.

P. Dostál, M. B?eznová, V. Kozlí?ková, T. Herben, and P. Ková?, Ant-induced soil modification and its effect on plant below-ground biomass, Pedobiologia, vol.49, pp.127-137, 2005.

J. Oliveras, J. M. Bas, and C. Gómez, Reduction of the ant mandible gap range after biotic homogenization caused by an ant invasion (Hymenoptera: Formicidae), Sociobiology, vol.45, pp.829-838, 2005.

J. Frouz, H. ?antr??ková, and J. Kal?ík, The effect of wood ants (Formica polyctena Foerst.) on the transformation of phosphorus in a spruce plantation, Pedobiologia, vol.41, pp.437-447, 1997.

D. Wagner and J. B. Jones, The impact of harvester ants on decomposition, N mineralization, litter quality, and the availability of N to plants in the Mojave Desert, Soil Biol. Biochem, vol.38, pp.2593-2601, 2006.

G. Brown, C. Scherber, P. Ramos, and E. K. Ebrahim, The effects of harvester ant (Messor ebeninus Forel) nests on vegetation and soil properties in a desert dwarf shrub community in north-eastern Arabia, Flora Morphol. Distrib. Funct. Ecol. Plants, vol.207, pp.503-511, 2012.

J. N. Nkem, L. De-bruyn, L. A. Grant, C. D. Hulugalle, and N. R. , The impact of ant bioturbation and foraging activities on surrounding soil properties, 2000.

, Pedobiologia, vol.44, p.70075

P. A. Domenico and F. W. Schwartz, Physical and chemical hydrogeology, 1998.

A. D. Davidson, J. K. Detling, and J. H. Brown, Ecological roles and conservation challenges of social, burrowing, herbivorous mammals in the world's grasslands, Front. Ecol. Environ, vol.10, pp.477-486, 2012.

B. Lafleur, L. M. Hooper-bùi, E. P. Mumma, and J. P. Geaghan, Soil fertility and plant growth in soils from pine forests and plantations: effect of invasive red imported fire ants Solenopsis invicta (Buren), Pedobiologia, vol.49, pp.415-423, 2005.

A. C. Risch, M. F. Jurgensen, M. Schütz, and D. S. Page-dumroese, The contribution of red wood ants to soil C and N pools and CO2 emissions in subalpine forests, Ecology, vol.86, pp.419-430, 2005.

M. Ohashi, L. Finér, T. Domisch, A. C. Risch, M. F. Jurgensen et al., Seasonal and diurnal CO2 efflux from red wood ant (Formica aquilonia) mounds in boreal coniferous forests, Soil Biol. Biochem, vol.39, pp.1504-1511, 2007.

M. Ohashi, J. Kilpeläinen, L. Finér, A. C. Risch, T. Domisch et al., The effect of red wood ant (Formica rufa group) mounds on root biomass, density, and nutrient concentrations in boreal managed forests, J. For. Res, vol.12, pp.113-119, 2007.

A. M. Boulton, B. A. Jaffee, and K. M. Scow, Effects of a common harvester ant (Messor andrei) on richness and abundance of soil biota, Appl. Soil Ecol, vol.23, pp.257-265, 2003.

A. Lang, J. Filser, and J. R. Henschel, Predation by ground beetles and wolf spiders on herbivorous insects in a maize crop, Agric. Ecosyst. Environ, vol.72, pp.189-199, 1999.

P. E. Cushing, Spider-ant associations: an updated review of myrmecomorphy, myrmecophily, and myrmecophagy in spiders, Psyche, p.151989, 2012.

T. Traxler, The impact of predation by the myrmecophagous spider Zodarion elegans (Araneae: Zodariidae) on the activity pattern of the Mediterranean harvester ant Messor wasmanni (Hymenoptera: Formicidae), Ecol. Montenegrina, vol.7, pp.328-344, 2016.

N. Gaur and B. P. Mohanty, Land-surface controls on near-surface soil moisture dynamics: traversing remote sensing footprints, Water Resour. Res, vol.52, pp.6365-6385, 2016.

S. K. Berke, Functional groups of ecosystem engineers: a proposed classification with comments on current issues, Integr. Comp. Biol, vol.50, pp.147-157, 2010.

L. R. Prugh and J. S. Brashares, Partitioning the effects of an ecosystem engineer: Kangaroo rats control community structure via multiple pathways, J. Anim. Ecol, vol.81, pp.667-678, 2012.

T. Liu, Earthworms coordinate soil biota to improve multiple ecosystem functions, Curr. Biol, vol.29, pp.3420-3429, 2019.

S. Matsuzaki, N. Usio, N. Takamura, and I. Washitani, Effects of common carp on nutrient dynamics and littoral community composition: roles of excretion and bioturbation, Fundam. Appl. Limnol, vol.168, pp.27-38, 2007.

N. Usio and C. R. Townsend, Roles of crayfish: consequences of predation and bioturbation for stream invertebrates, Ecology, vol.85, pp.807-822, 2004.