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TRY plant trait database – enhanced coverage and open access

Jens Kattge 1, 2, * Gerhard Bonisch 1 Sandra Díaz 3 Sandra Lavorel 4 Iain Colin Prentice 5 Paul Leadley 6 Susanne Tautenhahn 1 Gijsbert Werner 7, 8 Tuomas Aakala 9 Mehdi Abedi 10 Alicia Acosta 11 George Adamidis 12, 13 Kairi Adamson 14 Masahiro Aiba 15 Cécile Albert 16 Julio Alcántara 17 Carolina Alcázar C 18 Izabela Aleixo 19 Hamada Ali 20 Bernard Amiaud 21 Christian Ammer 22, 23 Mariano Amoroso 24, 25 Madhur Anand 26 Carolyn Anderson 27, 28 Niels Anten 29 Joseph Antos 30 Deborah Mattos Guimarães Apgaua 31 Tia‐lynn Ashman 32 Degi Harja Asmara 33 Gregory Asner 34 Michael Aspinwall 35 Owen Atkin 36 Isabelle Aubin 37 Lars Baastrup‐spohr 38 Khadijeh Bahalkeh 39 Michael Bahn 40 Timothy Baker 41 William Baker 42 Jan Bakker 43 Dennis Baldocchi 44 Jennifer Baltzer 45 Arindam Banerjee 46 Anne Baranger 47 Jos Barlow 48 Diego Barneche 49 Zdravko Baruch 50 Denis Bastianelli 51, 52 John Battles 53 William Bauerle 54 Marijn Bauters 55, 56 Erika Bazzato 57 Michael Beckmann 58 Hans Beeckman 59 Carl Beierkuhnlein 60 Renee Bekker 61 Gavin Belfry 62, 63 Michael Belluau 64 Mirela Beloiu 65 Raquel Benavides 66 Lahcen Benomar 67 Mary Lee Berdugo‐lattke 68, 69 Erika Berenguer 70 Rodrigo Bergamin 71 Joana Bergmann 72, 73 Marcos Bergmann Carlucci 74 Logan Berner 75 Markus Bernhardt‐römermann 76 Christof Bigler 77 Anne Bjorkman 78 Chris Blackman 79 Carolina Blanco 80 Benjamin Blonder 81, 62 Dana Blumenthal 82 Kelly Bocanegra‐gonzález 83 Pascal Boeckx 84 Stephanie Bohlman 85 Katrin Böhning‐gaese 86, 87 Laura Boisvert‐marsh 37 William Bond 88, 89 Ben Bond‐lamberty 90 Arnoud Boom 91 Coline Boonman 92 Kauane Bordin 93 Elizabeth Boughton 94 Vanessa Boukili 95 David Bowman 96 Sandra Bravo 97 Marco Richard Brendel 98 Martin Broadley 99 Kerry Brown 100 Helge Bruelheide 101, 2 Federico Brumnich 25, 102 Hans Henrik Bruun 38 David Bruy 103, 104 Serra Buchanan 105 Solveig Franziska Bucher 106 Nina Buchmann 77 Robert Buitenwerf 107, 108 Daniel Bunker 109 Jana Bürger 110 Sabina Burrascano 111 David Burslem 112 Bradley Butterfield 113 Chaeho Byun 114 Marcia Marques 115 Marina Scalon 116 Marco Caccianiga 117 Marc Cadotte 105 Maxime Cailleret 118, 119, 120 James Camac 121 Jesus Julio Camarero 122 Courtney Campany 123 Giandiego Campetella 124 Juan Antonio Campos 125 Laura Cano‐arboleda 126, 69 Roberto Canullo 124 Michele Carbognani 127 Fabio Carvalho 48 Fernando Casanoves 128 Bastien Castagneyrol 129 Jane Catford 130 Jeannine Cavender‐bares 131 Bruno Cerabolini 132 Marco Cervellini 124, 133 Eduardo Chacón‐madrigal 134 Kenneth Chapin 135 F. Stuart Chapin 136 Stefano Chelli 124 Si‐chong Chen 137 Anping Chen 138 Paolo Cherubini 139, 140 Francesco Chianucci 141 Brendan Choat 142 Kyong‐sook Chung 143 Milan Chytrý 144 Daniela Ciccarelli 145 Lluís Coll 146, 147 Courtney Collins 148 Luisa Conti 149, 150 David Coomes 151 Johannes Cornelissen 152 William Cornwell 153 Piermaria Corona 141 Marie Coyea 154 Joseph Craine 155 Dylan Craven 156 Joris Cromsigt 157, 158 Anikó Csecserits 159 Katarina Cufar 160 Matthias Cuntz 161 Ana Carolina Silva 162 Kyla Dahlin 163 Matteo Dainese 164 Igor Dalke 165 Michele Dalle Fratte 132 Anh Tuan Dang‐le 166 Jirí Danihelka 144, 150 Masako Dannoura 167, 168 Samantha Dawson 169 Arend Jacobus Beer 170 Angel de Frutos 2, 171 Jonathan de Long 172 Benjamin Dechant 173, 174, 175 Sylvain Delagrange 176, 177 Nicolas Delpierre 178 Géraldine Derroire 179 Arildo Dias 180 Milton Hugo Diaz‐toribio 181 Panayiotis Dimitrakopoulos 12 Mark Dobrowolski 182, 183 Daniel Doktor 58 Pavel Dřevojan 144 Ning Dong 184 John Dransfield 42 Stefan Dressler 185 Emilie Ducouret 179 Stefan Dullinger 186 Walter Durka 2, 58 Remko Duursma 142 Olga Dymova 165 Anna E‐vojtkó 150 Rolf Lutz Eckstein 187 Hamid Ejtehadi 188 James Elser 189, 190 Thaise Emilio 191 Kristine Engemann 107 Mohammad Bagher Erfanian 188 Alexandra Erfmeier 2, 192 Adriane Esquivel-Muelbert 41, 193 Gerd Esser 194 Marc Estiarte 195 Tomas Domingues 196 William Fagan 197 Jaime Fagúndez 198 Daniel Falster 199 Ying Fan 200 Jingyun Fang 201 Emmanuele Farris 202 Fatih Fazlioglu 203 Yanhao Feng 204 Fernando Fernandez‐mendez 205 Carlotta Ferrara 206 Joice Ferreira 207 Alessandra Fidelis 208 Bryan Finegan 209 Jennifer Firn 210 Timothy Flowers 211 Dan Flynn 212 Veronika Fontana 213 Estelle Forey 214 Cristiane Forgiarini 215 Louis François 216 Marcelo Frangipani 217 Dorothea Frank 1 Cedric Frenette‐dussault 218 Gregoire Freschet 219, 220 Ellen Fry 221 Nikolaos Fyllas 222 Guilherme Mazzochini 223 Sophie Gachet 16 Rachael Gallagher 224 Gislene Ganade 225 Francesca Ganga 226 Pablo García‐palacios 227 Veronica Gargaglione 228 Eric Garnier Jose Luis Garrido 229 André Luís Gasper 230 Guillermo Gea‐izquierdo 231 David Gibson 232 Andrew Gillison 233 Aelton Giroldo 234 Mary‐claire Glasenhardt 235 Sean Gleason 236 Mariana Gliesch 237 Emma Goldberg 238 Bastian Göldel 239 Erika Gonzalez‐akre 240 Jose Gonzalez‐andujar 241 Andrés González‐melo 242 Ana González‐robles 17 Bente Jessen Graae 243 Elena Granda 244 Sarah Graves 181 Walton Green 212 Thomas Gregor 245 Nicolas Gross 227 Greg Guerin 246 Angela Günther 1 Alvaro Gutiérrez 247 Lillie Haddock 248 Anna Haines 221 Jefferson Hall 249 Alain Hambuckers 250 Wenxuan Han 251 Sandy Harrison 252 Wesley Hattingh 253 Joseph Hawes 254 Tianhua He 255 Pengcheng He 251 Jacob Mason Heberling 256 Aveliina Helm 14 Stefan Hempel 72 Jörn Hentschel 106 Bruno Hérault 179 Ana‐maria Hereş 257 Katharina Herz 258 Myriam Heuertz 259 Thomas Hickler 260 Peter Hietz 261 Pedro Higuchi 262 Andrew Hipp 235 Andrew Hirons 263 Maria Hock 264 James Aaron Hogan 265 Karen Holl 266 Olivier Honnay 267 Daniel Hornstein 268 Enqing Hou 251 Nate Hough‐snee 269 Knut Anders Hovstad 270 Tomoaki Ichie 271 Boris Igić 272 Estela Illa 273 Marney Isaac 105 Masae Ishihara 274 Leonid Ivanov 275 Larissa Ivanova 165 Colleen Iversen 276 Jordi Izquierdo 277 Robert Jackson 278 Benjamin Jackson 279 Hervé Jactel 259 Andrzej Jagodzinski 280 Ute Jandt 281 Steven Jansen 282 Thomas Jenkins 283 Anke Jentsch 284 Jens Rasmus Plantener Jespersen 285 Guo‐feng Jiang 286 Jesper Liengaard Johansen 285 David Johnson 287 Eric Jokela 181 Carlos Alfredo Joly 223 Gregory Jordan 288 Grant Stuart Joseph 289 Decky Junaedi 290 Robert Junker 291 Eric Justes 292 Richard Kabzems 293 Jeffrey Kane 294 Zdenek Kaplan 295 Teja Kattenborn 296 Lyudmila Kavelenova 297 Elizabeth Kearsley 298 Anne Kempel 299 Tanaka Kenzo 300 Andrew Kerkhoff 301 Mohammed Khalil 302 Nicole Kinlock 303 Wilm Daniel Kissling 304 Kaoru Kitajima 265 Thomas Kitzberger 305 Rasmus Kjøller 306 Tamir Klein 307 Michael Kleyer 308 Jitka Klimešová 295 Joice Klipel 93 Brian Kloeppel 309 Stefan Klotz 58 Johannes Knops 310 Takashi Kohyama 311 Fumito Koike 312 Johannes Kollmann 313 Benjamin Komac 314 Kimberly Komatsu 315 Christian König 316 Nathan Kraft 317 Koen Kramer 318 Holger Kreft 319 Ingolf Kühn 58 Dushan Kumarathunge 142 Jonas Kuppler 320 Hiroko Kurokawa 15 Yoko Kurosawa 321 Shem Kuyah 322 Jean‐paul Laclau 323 Benoit Lafleur 324 Erik Lallai 226 Eric Lamb 325 Andrea Lamprecht 326 Daniel Larkin 238 Daniel Laughlin 327 Yoann Le Bagousse-Pinguet 16 Guerric Maire 323, 328 Peter Roux 170 Elizabeth Le Roux 329 Tali Lee 330 Frederic Lens 331 Simon Lewis 287 Barbara Lhotsky 155 Yuanzhi Li 332 Xine Li 333 Jeremy Lichstein 181 Mario Liebergesell 334 Jun Ying Lim 335 Yan‐shih Lin 224 Juan Carlos Linares 336 Chunjiang Liu 337 Daijun Liu 338 Udayangani Liu 339 Stuart Livingstone 105 Joan Llusia 195 Madelon Lohbeck 318 Gabriela Lopez‐gonzalez 340 Zdeňka Lososová 341 Frédérique Louault 342 Balázs Lukács 343 Petr Lukeš 344 Yunjian Luo 333 Michele Lussu 226 Siyan Ma 345 Camilla Maciel Rabelo Pereira 285 Michelle Mack 346 Vincent Maire 347 Annikki Mäkelä 9 Harri Mäkinen 348 Ana Claudia Mendes Malhado 349 Azim Mallik 350 Peter Manning 351 Stefano Manzoni 352 Zuleica Marchetti 353 Luca Marchino 206 Vinicius Marcilio‐silva 354 Eric Marcon 179 Michela Marignani 355 Lars Markesteijn 356 Adam Martin 105 Cristina Martínez‐garza 357 Jordi Martínez‐vilalta 358 Tereza Mašková 295 Kelly Mason 48 Norman Mason 359 Tara Joy Massad 360 Jacynthe Masse 361 Itay Mayrose 362 James Mccarthy 363 M. Luke Mccormack 238 Katherine Mcculloh 364 Ian Mcfadden 317 Brian Mcgill 365 Mara Mcpartland 238 Juliana Medeiros 366 Belinda Medlyn 367 Pierre Meerts 368 Zia Mehrabi 369 Patrick Meir 370 Felipe Melo 371 Maurizio Mencuccini 372 Céline Meredieu 259 Julie Messier 373 Ilona Meszaros 374 Juha Metsaranta 375 Sean Michaletz 369 Chrysanthi Michelaki 222 Svetlana Migalina 275 Rubén Milla 227 Jesse Miller 278 Vanessa Minden 376 Ray Ming 377 Karel Mokany 378 Angela Moles 379 Attila Molnár 380 Jane Molofsky 381 Martin Molz 382 Rebecca Montgomery 238 Arnaud Monty 383 Lenka Moravcová 384 Alvaro Moreno‐martínez 385 Marco Moretti 386 Akira Mori 312 Shigeta Mori 321 Dave Morris 387 Jane Morrison 277 Ladislav Mucina 255 Sandra Mueller 388 Christopher Muir 389 Sandra Cristina Muller 217 François Munoz 390 Isla Myers‐smith 279 Randall Myster 391 Masahiro Nagano 392 Shawna Naidu 377 Ayyappan Narayanan 390 Balachandran Natesan 390 Luka Negoita 393 Andrew Nelson 394 Eike Lena Neuschulz 260 Jian Ni 395 Georg Niedrist 396 Jhon Nieto 397 Rachael Nolan 142 Henning Nottebrock 398 Yann Nouvellon 323, 328 Alexander Novakovskiy 399 Kristin Odden Nystuen 243 Anthony O'Grady 400 Kevin O'Hara 53 Andrew O'Reilly‐nugent 401 Simon Oakley 402 Walter Oberhuber 403 Toshiyuki Ohtsuka 404 Ricardo Oliveira 405 Mark Olson 406 Vladimir Onipchenko 407 Yusuke Onoda 274 Renske Onstein 2 Jenny Ordoñez 408 Noriyuki Osada 409 Ivika Ostonen 410 Gianluigi Ottaviani 411 Sarah Otto 369 Gerhard Overbeck 217 Wim Ozinga 412 Anna Pahl 313 C. Paine 413 Robin Pakeman 414 Aristotelis Papageorgiou 415 Evgeniya Parfionova 297 Meelis Pärtel 416 Marco Patacca 318 Susana Paula 417 Juraj Paule 418 Harald Pauli 419 Juli Pausas 420 Begoña Peco 421 Josep Peñuelas 195 Antonio Perea 422 Pablo Luis Peri 228 Ana Carolina Petisco‐souza 354 Alessandro Petraglia 423 Any Mary Petritan 424 Oliver Phillips 340 Simon Pierce 425 Valério Pillar 225 Jan Pisek 426 Alexandr Pomogaybin 427 Hendrik Poorter 428 Angelika Portsmuth 429 Peter Poschlod 430 Catherine Potvin 249 Devon Pounds 235 A. Shafer Powell 276 Sally Power 431 Andreas Prinzing 283 Giacomo Puglielli 432 Petr Pyšek 384 Valérie Raevel 390 Anja Rammig 433 Johannes Ransijn 285 Courtenay Ray 434 Peter Reich 435 Markus Reichstein 1 Douglas Reid 436 Maxime Réjou-Méchain 103 Victor Resco Dios 437 Sabina Ribeiro 438 Sarah Richardson 359 Kersti Riibak 14 Matthias Rillig 439 Fiamma Riviera 182 Elisabeth Robert 440 Scott Roberts 441 Bjorn Robroek 442 Adam Roddy 443 Arthur Vinicius Rodrigues 217 Alistair Rogers 444 Emily Rollinson 445 Victor Rolo 446 Christine Romermann 447 Dina Ronzhina 165 Christiane Roscher 2 Julieta Rosell 406 Milena Fermina Rosenfield 448 Christian Rossi 449 David Roy 450 Samuel Royer‐tardif 451 Nadja Rüger 2 Ricardo Ruiz‐peinado 452 Sabine Rumpf 419 Graciela Rusch 453 Masahiro Ryo 72 Lawren Sack 317 Angela Saldaña 406 Beatriz Salgado‐negret 454 Roberto Salguero‐gomez 455 Ignacio Santa‐regina 456 Ana Carolina Santacruz‐garcía 97 Joaquim Santos 457 Jordi Sardans 195 Brandon Schamp 458 Michael Scherer‐lorenzen 388 Matthias Schleuning 260 Bernhard Schmid 449 Marco Schmidt 459 Sylvain Schmitt 259 Julio Schneider 245 Simon Schowanek 239 Julian Schrader 460 Franziska Schrodt 461 Bernhard Schuldt 462 Frank Schurr 463 Galia Selaya Garvizu 464 Marina Semchenko 221 Colleen Seymour 465 Julia Sfair 466 Joanne Sharpe 467 Christine Sheppard 468 Serge Sheremetiev 469 Satomi Shiodera 470 Bill Shipley 471 Tanvir Ahmed Shovon 472 Alrun Siebenkäs 473 Carlos Sierra 1 Vasco Silva 474 Mateus Silva 475 Tommaso Sitzia 476 Henrik Sjöman 477 Martijn Slot 478 Nicholas Smith 479 Darwin Sodhi 369 Pamela Soltis 480 Douglas Soltis 181 Ben Somers 481 Grégory Sonnier 179 Mia Vedel Sørensen 243 Enio Egon Sosinski 482 Nadejda Soudzilovskaia 483 Alexandre Souza 225 Marko Spasojevic 484 Marta Gaia Sperandii 11 Amanda Stan 346 James Stegen 485 Klaus Steinbauer 486 Jörg Stephan 487 Frank Sterck 318 Dejan Stojanovic 488 Tanya Strydom 352 Maria Laura Suarez 489 Jens‐christian Svenning 239 Ivana Svitková 490 Marek Svitok 491 Miroslav Svoboda 492 Emily Swaine 493 Nathan Swenson 494 Marcelo Tabarelli 371 Kentaro Takagi 311 Ulrike Tappeiner 495 Rubén Tarifa 496 Simon Taugourdeau 51 Cagatay Tavsanoglu 497 Mariska Te Beest 329 Leho Tedersoo 14 Nelson Thiffault 498 Dominik Thom 381 Evert Thomas 499 Ken Thompson 500 Peter Thornton 276 Wilfried Thuiller 4 Lubomír Tichý 295 David Tissue 142 Mark Tjoelker 142 David Yue Phin Tng 501 Joseph Tobias 5 Peter Török 502 Tonantzin Tarin 503 Jose Manuel Torres Ruiz 504 Béla Tóthmérész 505 Martina Treurnicht 506 Valeria Trivellone 377 Franck Trolliet 383 Volodymyr Trotsiuk 507 James Tsakalos 182 Ioannis Tsiripidis 508 Niklas Tysklind 179 Toru Umehara 509 Vladimir Usoltsev 510 Matthew Vadeboncoeur 511 Jamil Vaezi 512 Fernando Valladares 513 Jana Vamosi 514 Peter van Bodegom 483 Michiel Breugel 443 Elisa van Cleemput 515 Martine Weg Stephni Merwe Fons Plas Masha Sande Mark Kleunen Koenraad van Meerbeek Mark Vanderwel Kim André Vanselow Angelica Varhammar 516 Laura Varone Maribel Yesenia Vasquez Valderrama Kiril Vassilev Mark Vellend 369 Erik Veneklaas Hans Verbeeck 517 Kris Verheyen 518 Alexander Vibrans Ima Vieira 519 Jaime Villacís Cyrille Violle 220 Pandi Vivek Katrin Wagner Matthew Waldram Anthony Waldron Anthony Walker Martyn Waller Gabriel Walther Han Wang 520 Feng Wang Weiqi Wang 521 Harry Watkins James Watkins Ulrich Weber James Weedon 522 Liping Wei 523 Patrick Weigelt 319 Evan Weiher 524 Aidan Wells Camilla Wellstein 525 Elizabeth Wenk Mark Westoby 526 Alana Westwood Philip John White Mark Whitten Mathew Williams 372 Daniel Winkler 266 Klaus Winter 249 Chevonne Womack 170 Ian Wright 224 S. Joseph Wright 249 Justin Wright 527 Bruno Pinho 528 Fabiano Ximenes 529 Toshihiro Yamada 530 Keiko Yamaji 531 Ruth Yanai 532 Nikolay Yankov 427 Benjamin Yguel 283 Kátia Janaina Zanini 93 Amy Zanne 533 David Zelený 534 Yun‐peng Zhao 535 Jingming Zheng 536 Ji Zheng 337 Kasia Ziemińska 5 Chad Zirbel 238 Georg Zizka 180 Irié Casimir Zo‐bi 537 Gerhard Zotz 376 Christian Wirth 538
* Corresponding author
402 CEH
404 Department of Systems and Science
Graduate School of Informatics